title: List of ground beetle genera

abstract: This page lists the genera in the family Carabidae, according to Catalogue of Life and Carabcat Database as of March 2023, excluding Tiger Beetles.

text:
# List of ground beetle genera

This page lists the genera in the family Carabidae, according to Catalogue of Life and Carabcat Database as of March 2023, excluding Tiger Beetles (which are considered to be in the family Cicindelidae).

## Family Carabidae

### Subfamily Anthiinae Bonelli, 1813

Tribe Anthiini Bonelli, 1813 *Anthia* Weber, 1801 *Atractonotus* Perroud, 1847 *Baeoglossa* Chaudoir, 1850 *Cycloloba* Chaudoir, 1850 *Cypholoba* Chaudoir, 1850 *Eccoptoptera* Chaudoir, 1878 *Gonogenia* Chaudoir, 1844 *Netrodera* Chaudoir, 1850 Tribe Helluonini Hope, 1838 Subtribe Helluonina Hope, 1838 *Aenigma* Newman, 1836 *Ametroglossus* Sloane, 1914 *Dicranoglossus* Chaudoir, 1872 *Epimicodema* Sloane, 1914 *Gigadema* J.Thomson, 1859 *Helluapterus* Sloane, 1914 *Helluarchus* Sloane, 1914 *Helluo* Bonelli, 1813 *Helluodema* Laporte, 1867 *Helluonidius* Chaudoir, 1872 *Helluopapua* Darlington, 1968 *Helluosoma* Laporte, 1867 *Neohelluo* Sloane, 1914 *Platyhelluo* Baehr, 2005 Subtribe Omphrina Jedlicka, 1941 *Colfax* Andrewes, 1920 *Creagris* Nietner, 1857 *Dailodontus* Reiche, 1843 *Erephognathus* Alluaud, 1932 *Helluobrochus* Reichardt, 1974 *Helluomorpha* Laporte, 1834 *Helluomorphoides* Ball, 1951 *Macrocheilus* Hope, 1838 *Meladroma* Motschulsky, 1855 *Omphra* Dejean, 1825 *Pleuracanthus* Gray, 1832 *Triaenogenius* Chaudoir, 1877 Tribe Physocrotaphini Chaudoir, 1863 *Anguloderus* Anichtchenko & Sciaky, 2017 *Foveocrotaphus* Anichtchenko, 2014 *Helluodes* Westwood, 1847 *Holoponerus* Fairmaire, 1883 *Physocrotaphus* Parry, 1849 *Physoglossus* Akhil & Sabu, 2020 *Pogonoglossus* Chaudoir, 1863 *Schuelea* Baehr, 2004

### Subfamily Apotominae LeConte, 1853

*Apotomus* Illiger, 1807

### Subfamily Brachininae Bonelli, 1810

Tribe Brachinini Bonelli, 1810 *Aptinoderus* Hubenthal, 1919 *Aptinus* Bonelli, 1810 *Brachinulus* Basilewsky, 1958 *Brachinus* Weber, 1801 *Mastax* Fischer von Waldheim, 1828 *Pheropsophus* Solier, 1833 *Styphlodromus* Basilewsky, 1959 *Styphlomerus* Chaudoir in Putzeys, 1875 Tribe Crepidogastrini Jeannel, 1949 *Brachynillus* Reitter, 1904 *Crepidogaster* Boheman, 1848 *Crepidogastrillus* Basilewsky, 1959 *Crepidogastrinus* Basilewsky, 1957 *Crepidolomus* Basilewsky, 1959 *Crepidonellus* Basilewsky, 1959

### Subfamily Broscinae Hope, 1838

Tribe Broscini Hope, 1838 Subtribe Axonyina Roig-Juñent, 2000 *Broscodes* Bolivar y Pieltain, 1914 *Rawlinsius* Davidson & Ball, 1998 Subtribe Baripodina Jeannel, 1941 *Baripus* Dejean, 1828 *Bembidiomorphum* Champion, 1918 Subtribe Broscina Hope, 1838 *Broscodera* Lindroth, 1961 *Broscosoma* Rosenhauer, 1846 *Broscus* Panzer, 1813 *Chaetobroscus* Semenov, 1900 *Craspedonotus* Schaum, 1863 *Eobroscus* Kryzhanovskij, 1951 *Kashmirobroscus* J.Schmidt; Wrase & Sciaky, 2013 *Miscodera* Eschscholtz, 1830 *Zacotus* LeConte, 1869 Subtribe Creobiina Jeannel, 1941 *Acallistus* Sharp, 1886 *Adotela* Laporte, 1867 *Anheterus* Putzeys, 1868 *Bountya* Townsend, 1971 *Brithysternum* W.J.MacLeay, 1873 *Cascellius* Curtis, 1838 *Cerotalis* Laporte, 1867 *Creobius* Guérin-Méneville, 1838 *Gnathoxys* Westwood, 1842 *Nothocascellius* Roig-Juñent, 1995 *Promecoderus* Dejean, 1829 Subtribe Nothobroscina Roig-Juñent, 2000 *Chylnus* Sloane, 1920 *Diglymma* Sharp, 1886 *Eurylychnus* Bates, 1891 *Mecodema* Blanchard, 1843 *Monteremita* Seldon & Holwell, 2019 *Nothobroscus* Roig-Juñent & Ball, 1995 *Oregus* Putzeys, 1868 *Orthoglymma* Liebherr; Marris; Emberson; Syrett & Roig-Juñent, 2011 *Percolestus* Sloane, 1892 *Percosoma* Schaum, 1858

### Subfamily Carabinae Linnaeus, 1802

Tribe Carabini Linnaeus, 1802 Subtribe Carabina Linnaeus, 1802 *Aplothorax* G.R.Waterhouse, 1842 *Calosoma* Weber, 1801 *Carabus* Linnaeus, 1758 Subtribe Ceroglossina Lapouge, 1927 *Ceroglossus* Solier, 1848 Tribe Cychrini Perty, 1830 Subtribe Cychrina Perty, 1830 *Cychropsis* Boileau, 1901 *Cychrus* Fabricius, 1794 *Scaphinotus* Dejean, 1826 *Sphaeroderus* Dejean, 1826 Subtribe Pamborina Hope, 1838 *Maoripamborus* Brookes, 1944 *Pamborus* Latreille, 1812

### Subfamily Ctenodactylinae Laporte, 1834

Tribe Ctenodactylini Laporte, 1834 *Alachnothorax* Liebke, 1929 *Amblycoleus* Chaudoir, 1872 *Antipionycha* Liebke, 1928 *Askalaphium* Liebke, 1938 *Ctenodactyla* Dejean, 1825 *Leptotrachelon* Liebke, 1928 *Leptotrachelus* Latreille, 1829 *Oilea* Liebke, 1931 *Parapionycha* Liebke, 1929 *Pionycha* Chaudoir, 1848 *Plagiotelum* Solier, 1849 *Propionycha* Liebke, 1928 *Pseudometabletus* Liebke, 1930 *Schidonychus* Klug, 1834 *Teukrus* Liebke, 1931 *Wate* Liebke, 1928 Tribe Hexagoniini G.Horn, 1881 *Dinopelma* Bates, 1889 *Hexagonia* Kirby, 1825 *Omphreoides* Fairmaire, 1896

### Subfamily Dryptinae Bonelli, 1810

Tribe Dryptini Bonelli, 1810 *Desera* Hope, 1831 *Drypta* Latreille, 1797 *Megadrypta* Sciaky & Anichtchenko, 2020 *Neodrypta* Basilewsky, 1960 *Nesiodrypta* Jeannel, 1949 *Prionodrypta* Jeannel, 1949 Tribe Galeritini Kirby, 1825 *Ancystroglossus* Chaudoir, 1863 *Eunostus* Laporte, 1835 *Galerita* Fabricius, 1801 *Trichognatha* Latreille, 1829 Tribe Zuphiini Bonelli, 1810 Subtribe Dicrodontina Machado, 1992 *Dicrodontus* Chaudoir, 1872 Subtribe Leleupidiina Basilewsky, 1951 *Colasidia* Basilewsky, 1954 *Gunvorita* Landin, 1955 *Leleupidia* Basilewsky, 1951 *Neoleleupidia* Basilewsky, 1953 *Paraleleupidia* Basilewsky, 1951 Subtribe Planetina Jedlicka, 1941 *Planetes* W.S.MacLeay, 1825 Subtribe Zuphiina Bonelli, 1810 *Acrogenys* W.J.MacLeay, 1864 *Agastus* Schmidt-Goebel, 1846 *Chaudoirella* Mateu, 1982 *Coarazuphium* Gnaspini; Vanin & Godoy, 1998 *Ildobates* Español, 1966 *Metaxidius* Chaudoir, 1852 *Metazuphium* Mateu, 1992 *Mischocephalus* Chaudoir, 1863 *Parazuphium* Jeannel, 1942 *Polistichus* Bonelli, 1810 *Pseudaptinus* Laporte, 1834 *Speothalpius* B.Moore, 1995 *Speozuphium* B.Moore, 1995 *Typhlozuphium* Baehr, 2014 *Zuphioides* Ball & Shpeley, 2013 *Zuphium* Latreille, 1805

### Subfamily Elaphrinae Latreille, 1802

*Blethisa* Bonelli, 1810 *Diacheila* Motschulsky, 1844 *Elaphrus* Fabricius, 1775 *Elaphrotites* Haupt, 1856

### Subfamily Gineminae Ball & Shpeley, 2002

*Ginema* Ball & Shpeley, 2002

### Subfamily Harpalinae Bonelli, 1810

Tribe Anisodactylini Lacordaire, 1854 *Allendia* Noonan, 1974 *Allocinopus* Broun, 1903 *Amphasia* Newman, 1838 *Anisodactylus* Dejean, 1829 *Anisostichus* Emden, 1953 *Cenogmus* Sloane, 1898 *Chydaeus* Chaudoir, 1854 *Crasodactylus* Guérin-Méneville, 1847 *Criniventer* Emden, 1953 *Diachromus* Erichson, 1837 *Dicheirus* Mannerheim, 1843 *Gaioxenus* Broun, 1910 *Geopinus* LeConte, 1847 *Gnathaphanus* W.S.MacLeay, 1825 *Gynandromorphus* Dejean, 1829 *Haplaner* Chaudoir, 1878 *Harpalomimetes* Schauberger in Csiki, 1932 *Hypharpax* W.S.MacLeay, 1825 *Maoriharpalus* Larochelle & Larivière, 2005 *Nornalupia* Kataev, 2002 *Notiobia* Perty, 1830 *Parabaris* Broun, 1881 *Progonochaetus* G.Müller, 1938 *Pseudanisotarsus* Noonan, 1973 *Pseudognathaphanus* Schauberger, 1932 *Pseudorhysopus* Kataev & Wrase, 2001 *Rhysopus* Andrewes, 1929 *Scybalicus* Schaum, 1862 *Triplosarus* Bates, 1874 *Tuiharpalus* Larochelle & Larivière, 2005 *Xestonotus* LeConte, 1853 Tribe Harpalini Bonelli, 1810 Subtribe Amblystomina Fauvel, 1889 *Amblystomus* Erichson, 1837 *Anomostomus* LaFerté-Sénectère, 1853 *Barysomus* Dejean, 1829 *Oosoma* Nietner, 1857 Subtribe Ditomina Bonelli, 1810 *Bronislavia* Semenov, 1891 *Carenochyrus* Solsky, 1874 *Carterus* Dejean, 1830 *Chilotomus* Chaudoir, 1842 *Ditomus* Bonelli, 1810 *Dixus* Billberg, 1820 *Eocarterus* Stichel, 1923 *Eucarterus* Reitter, 1900 *Graniger* Motschulsky, 1864 *Indocarterus* Kataev & Wrase, 2018 *Liochirus* Tschitscherine, 1897 *Machozetus* Chaudoir, 1850 *Odotoncarus* Solier, 1835 *Oedesis* Motschulsky, 1850 *Pachycarus* Solier, 1835 *Parapenthes* Kataev & Wrase, 2018 *Penthus* Chaudoir, 1843 *Phorticosomus* Schaum, 1863 *Proditomus* Schauberger, 1934 *Pseudaristus* Reitter, 1900 *Tschitscherinellus* Csiki, 1906 Subtribe Harpalina Bonelli, 1810 *Acinopus* Dejean, 1821 *Afromizonus* Basilewsky, 1947 *Allosiopelus* N.Ito, 1995 *Amblygnathus* Dejean, 1829 *Anisocnemus* Chaudoir, 1843 *Athrostictus* Bates, 1878 *Axinotoma* Dejean, 1829 *Aztecarpalus* Ball, 1970 *Bleusea* Bedel, 1897 *Boeomimetes* Péringuey, 1896 *Bradybaenus* Dejean, 1829 *Coleolissus* Bates, 1892 *Cratacanthus* Dejean, 1829 *Cratognathus* Dejean, 1829 *Daptus* Fischer von Waldheim, 1823 *Dioryche* W.S.MacLeay, 1825 *Discoderus* LeConte, 1853 *Dregus* Motschulsky, 1864 *Ectinothorax* Alluaud, 1941 *Eriophonus* Tschitscherine, 1901 *Euryderus* LeConte, 1846 *Geodromus* Dejean, 1829 *Harpalinus* Jeannel, 1946 *Harpaliscus* Bates, 1892 *Harpalobrachys* Tschitscherine, 1899 *Harpalodiodes* Bousquet, 2002 *Harpalomimus* Facchini & Giachino, 2020 *Harpalomorphus* Péringuey, 1896 *Harpalus* Latreille, 1802 *Harpathaumas* Basilewsky, 1947 *Harponixus* Basilewsky, 1950 *Hartonymus* Casey, 1914 *Heteracantha* Brullé, 1835 *Hyphaereon* W.S.MacLeay, 1825 *Indiophonus* N.Ito, 1996 *Kareya* Andrewes, 1919 *Lampetes* Andrewes, 1940 *Laparhetes* Jeannel, 1946 *Liodaptus* Bates, 1889 *Meroctenus* Gemminger & Harold, 1868 *Microderes* Faldermann, 1836 *Neoaulacoryssus* Noonan, 1985 *Neodiachipteryx* Noonan, 1985 *Neohyparpalus* Clarke, 1981 *Neophygas* Noonan, 1976 *Nesacinopus* Tschitscherine, 1900 *Nesarpalus* Bedel, 1897 *Nipponoharpalus* Habu, 1973 *Nothodaptus* Maindron, 1906 *Oesyperus* Andrewes, 1923 *Omostropus* Péringuey, 1896 *Ooidius* Chaudoir, 1847 *Ophoniscus* Bates, 1892 *Ophonus* Dejean, 1821 *Oxycentrus* Chaudoir, 1854 *Panagrius* Andrewes, 1933 *Pangus* Dejean, 1821 *Parasiopelus* Basilewsky, 1946 *Paraulacoryssus* Shpeley; Hunting & Ball, 2017 *Parophonus* Ganglbauer, 1891 *Paulianoscirtus* Basilewsky, 1976 *Penthophonus* Reitter, 1900 *Piosoma* LeConte, 1847 *Platymetopsis* Ball & Maddison, 1987 *Platymetopus* Dejean, 1829 *Prakasha* Andrewes, 1919 *Pseudodiachipteryx* Burgeon, 1936 *Pseudohyparpalus* Basilewsky, 1946 *Pseudoselenophorus* Péringuey, 1896 *Selenophorus* Dejean, 1829 *Selenotichnus* Kataev, 1999 *Siopelophonus* Kataev, 2015 *Siopelus* Murray, 1859 *Stenomorphus* Dejean, 1831 *Trichopselaphus* Chaudoir, 1843 *Trichotichnus* A.Morawitz, 1863 *Trichoxycentrus* N.Ito, 2000 *Typsiharpalus* Tschitscherine, 1901 Tribe Pelmatellini Bates, 1882 *Hakaharpalus* Larochelle & Larivière, 2005 *Kupeharpalus* Larochelle & Larivière, 2005 *Lecanomerus* Chaudoir, 1850 *Nemaglossa* Solier, 1849 *Notospeophonus* B.Moore, 1962 *Pelmatellus* Bates, 1882 *Syllectus* Bates, 1878 *Trachysarus* Reed, 1874 Tribe Stenolophini Kirby, 1837 *Acupalpus* Latreille, 1829 *Agonoleptus* Casey, 1914 *Amerinus* Casey, 1884 *Angionychus* Klug, 1853 *Anthracus* Motschulsky, 1850 *Batoscelis* Dejean, 1836 *Bradycellus* Erichson, 1837 *Cratosoma* Jeannel, 1948 *Cyptomicrus* Vinson, 1939 *Dicheirotrichus* Jacquelin du Val, 1855 *Egadyla* Alluaud, 1916 *Euthenarus* Bates, 1874 *Fuminoria* Morita, 2006 *Goniocellus* Casey, 1914 *Gugheorites* Basilewsky, 1951 *Haplanister* B.Moore, 1996 *Hemiaulax* Bates, 1892 *Hippoloetis* Laporte, 1835 *Idiomelas* Tschitscherine, 1900 *Kaffovatus* Clarke, 1972 *Kenyacus* Alluaud, 1917 *Kiwiharpalus* Larochelle & Larivière, 2005 *Lioholus* Tschitscherine, 1897 *Loxoncus* Schmidt-Goebel, 1846 *Pachytrachelus* Chaudoir, 1852 *Parabradycellus* N.Ito, 2003 *Paramecus* Dejean, 1829 *Philodes* LeConte, 1861 *Pholeodytes* Britton, 1962 *Pogonodaptus* G.Horn, 1881 *Polpochila* Solier, 1849 *Psychristus* Andrewes, 1930 *Rhabidius* Basilewsky, 1948 *Stenolophus* Dejean, 1821 *Uenanthracus* Kasahara, 1994

### Subfamily Hiletinae Schiödte, 1847

*Eucamaragnathus* Jeannel, 1938 *Hiletus* Schiödte, 1847

### Subfamily Lebiinae Bonelli, 1810

Tribe Cyclosomini Laporte, 1834 Subtribe Corsyrina Ganglbauer *Corsyra* Dejean, 1825 *Discoptera* Semenov, 1889 Subtribe Cyclosomina Laporte, 1834 *Cyclosomus* Latreille, 1829 *Sarothrocrepis* Chaudoir, 1850 *Tetragonoderus* Dejean, 1829 Subtribe Graphipterina Latreille, 1802 *Graphipterus* Latreille, 1802 *Piezia* Brullé, 1834 *Trichopiezia* Nègre, 1955 Subtribe Masoreina Chaudoir, 1871 *Anaulacus* W.S.MacLeay, 1825 *Atlantomasoreus* Mateu, 1984 *Leuropus* Andrewes, 1947 *Lophidius* Dejean, 1831 *Masoreus* Dejean, 1821 *Mnuphorus* Chaudoir, 1873 *Odontomasoreus* Darlington, 1968 *Somoplatodes* Basilewsky, 1986 *Somoplatus* Dejean, 1829 Subtribe Nemotarsina Bates, 1883 *Nemotarsus* LeConte, 1853 Tribe Lachnophorini LeConte, 1853 Subtribe Calophaenina Jeannel, 1948 *Calophaena* Klug, 1821 *Calophaenoidea* Liebke, 1930 Subtribe Lachnophorina LeConte, 1853 *Amphithasus* Bates, 1871 *Anchonoderus* Reiche, 1843 *Aporesthus* Bates, 1871 *Asklepia* Liebke, 1938 *Balligratus* Moret & Ortuño, 2017 *Calybe* Laporte, 1834 *Ega* Laporte, 1834 *Eucaerus* LeConte, 1853 *Euphorticus* G.Horn, 1881 *Grundmannius* Basilewsky, 1965 *Guatemalteca* Erwin, 2004 *Lachnaces* Bates, 1872 *Lachnophorus* Dejean, 1831 *Peruphorticus* Erwin & Zamorano, 2014 *Pseudophorticus* Erwin, 2004 *Selina* Motschulsky, 1858 Tribe Lebiini Bonelli, 1810 Subtribe Agrina Kirby, 1837 *Abaditicus* Ball & Hilchie, 1983 *Abrodiella* Bousquet, 2002 *Agra* Fabricius, 1801 *Allardina* Basilewsky, 1963 *Amelus* Chaudoir, 1873 *Anomotariella* Baehr, 2012 *Anomotarus* Chaudoir, 1875 *Apterodromites* Mateu, 1976 *Aspasiola* Chaudoir, 1877 *Calleida* Latreille, 1824 *Calleidomorpha* Motschulsky, 1855 *Callidadelpha* Steinheil, 1875 *Chaudoirina* Mateu, 1955 *Cryptobatis* Eschscholtz, 1829 *Cyanotarus* Reed, 1874 *Cylindronotum* Putzeys, 1845 *Demetrida* White, 1846 *Diabaticus* Bates, 1878 *Do* Baehr, 2009 *Dromidea* Perroud & Montrouzier, 1864 *Epikastea* Liebke, 1935 *Eujalmenus* Bousquet, 2002 *Euplatia* Chaudoir, 1873 *Eurycalleida* Maindron, 1905 *Falsodromius* Mateu, 1976 *Glycia* Chaudoir, 1842 *Hyboptera* Chaudoir, 1873 *Hybopteroides* Erwin & Ball, 2012 *Infernophilus* Larson, 1969 *Kteatus* Liebke, 1936 *Lipostratia* Chaudoir, 1873 *Merizomena* Chaudoir, 1873 *Mimodromius* Chaudoir, 1873 *Mimophilorhizus* Mateu, 1993 *Oechalius* Liebke, 1935 *Ogygium* Liebke, 1935 *Onota* Chaudoir, 1873 *Otoglossa* Chaudoir, 1873 *Paraglycia* Bedel, 1904 *Phacocerus* Chaudoir, 1873 *Philophuga* Motschulsky, 1859 *Phloeocarabus* W.J.MacLeay, 1871 *Plochionus* Dejean, 1821 *Pontonoa* Liebke, 1935 *Pseudotoglossa* Mateu, 1961 *Pylartesius* Liebke, 1939 *Rugitarus* Baehr, 2009 *Speotarus* B.Moore, 1964 *Straneotia* Mateu, 1961 *Tecnophilus* Chaudoir, 1877 *Teiresia* Liebke, 1935 *Thoasia* Liebke, 1939 *Titaresius* Liebke, 1935 *Trigonothops* W.J.MacLeay, 1864 *Valeriaaschero* Erwin, 2004 *Vianasia* Mateu, 1955 Subtribe Apenina Ball *Apenes* LeConte, 1851 *Cymindoidea* Laporte, 1833 *Habutarus* Ball & Hilchie, 1983 *Platytarus* Fairmaire, 1850 *Trymosternus* Chaudoir, 1873 Subtribe Celaenephina Habu, 1982 *Celaenephes* Schmidt-Goebel, 1846 Subtribe Cymindidina Laporte, 1834 *Afrotarus* Jeannel, 1949 *Assadecma* Basilewsky, 1982 *Cymindis* Latreille, 1805 *Hystrichopus* Boheman, 1848 *Leptosarcus* Péringuey, 1896 *Metaxymorphus* Chaudoir, 1850 *Petrimagnia* Kryzhanovskij & Mikhailov, 1971 *Pinacodera* Schaum, 1857 *Plagiopyga* Boheman, 1848 *Pseudomasoreus* Desbrochers des Loges, 1904 *Taridius* Chaudoir, 1875 Subtribe Demetriadina Bates, 1886 *Cylindrocranius* Chaudoir, 1878 *Demetrias* Bonelli, 1810 Subtribe Dromiusina Bonelli, 1810 *Afrodromius* Basilewsky, 1958 *Axinopalpus* LeConte, 1846 *Barrymooreana* Baehr, 1997 *Brachynopterus* Bedel, 1898 *Brigalowia* Baehr, 2006 *Calodromius* Reitter, 1905 *Carbonellia* Mateu, 1968 *Disciferella* Kataev & Muilwijk, 2020 *Dromiops* Péringuey, 1898 *Dromius* Bonelli, 1810 *Dromoceryx* Schmidt-Goebel, 1846 *Geoffreyella* Baehr, 2012 *Klepteromimus* Péringuey, 1898 *Mesolestes* Schatzmayr, 1943 *Metadromius* Bedel, 1907 *Microdaccus* Schaum in Baudi di Selve, 1864 *Microlestes* Schmidt-Goebel, 1846 *Microlestodes* Baehr, 1987 *Monnea* Mateu, 1970 *Negrea* Mateu, 1968 *Omophagus* Andrewes, 1937 *Oxoides* Solier, 1849 *Paradromius* Fowler, 1887 *Philorhizus* Hope, 1838 *Polyaulacus* Chaudoir, 1878 *Psammodromius* Peyerimhoff, 1927 *Pseudomonnea* Mateu, 1983 *Somalodromius* Mateu, 1967 *Xenodromius* Bates, 1891 *Zolotarevskyella* Mateu, 1953 Subtribe Gallerucidiina Chaudoir, 1872 *Gallerucidia* Chaudoir, 1872 *Lebidia* A.Morawitz, 1862 Subtribe Lebiina Bonelli, 1810 *Alkestis* Liebke, 1939 *Aristolebia* Bates, 1892 *Daer* Semenov & Znojko, 1929 *Lachnolebia* Maindron, 1905 *Lebia* Latreille, 1802 *Lebidema* Motschulsky, 1864 *Lebiomorphica* Lorenz, 1998 *Lebistina* Motschulsky, 1864 *Lebistinida* Péringuey, 1898 *Lionedya* Chaudoir, 1871 *Matabele* Péringuey, 1896 *Megalebia* Mateu, 1972 *Orthobasis* Chaudoir, 1871 *Pachylebia* Jeannel, 1949 *Pachylebiodes* Mateu, 1972 *Paulianolebia* Mateu, 1972 *Pseudopachylebia* Mateu, 1972 *Rhopalostyla* Chaudoir, 1850 *Scalidion* Schmidt-Goebel, 1846 *Setolebia* Jedlicka, 1941 *Sofota* Jedlicka, 1951 Subtribe Lichnasthenina J.Thomson *Astastus* Péringuey, 1896 *Australovelinda* Baehr, 2012 *Imasakaia* Arai & Morita, 2010 *Lichnasthenus* J.Thomson, 1858 *Mimovelindopsis* Mateu, 1963 *Velinda* Andrewes, 1921 *Velindastus* Schüle & Lorenz, 2008 *Velindomimus* Jeannel, 1955 *Velindopsis* Burgeon, 1937 Subtribe Lionychina Jeannel, 1948 *Apristomimus* Mateu, 1969 *Apristus* Chaudoir, 1846 *Eremolestes* Maindron, 1905 *Lionychus* Wissmann, 1846 *Lorestania* Anichtchenko, 2011 *Metablus* Jedlicka, 1958 *Omobrus* Andrewes, 1930 *Singiliomimus* Péringuey, 1896 *Syntomus* Hope, 1838 *Tilius* Chaudoir, 1876 *Trichidema* Basilewsky, 1956 Subtribe Metallicina Basilewsky, 1984 *Euproctinus* Leng & Mutchler, 1927 *Metallica* Chaudoir, 1873 *Pachycallida* Jeannel, 1949 *Parena* Motschulsky, 1860 Subtribe Peliocypadina Basilewsky, 1984 *Demetriola* Jeannel, 1949 *Peliocypas* Schmidt-Goebel, 1846 Subtribe Pericalina Hope, 1838 *Agonocheila* Chaudoir, 1848 *Allophanes* Andrewes, 1939 *Allophanopsis* Louwerens, 1952 *Amphimenes* Bates, 1873 *Amphimenoides* Kirschenhofer, 1999 *Antimerina* Alluaud, 1897 *Arsinoe* Laporte, 1834 *Auchmerus* Andrewes, 1929 *Bellavalentis* Hunting & Yang, 2019 *Brachichila* Chaudoir, 1870 *Brachyctis* Chaudoir, 1870 *Catascopellus* Straneo, 1969 *Catascopus* Kirby, 1825 *Cheilagona* Baehr, 2006 *Coptodera* Dejean, 1825 *Coptoglossus* Chaudoir, 1870 *Coptoptera* Chaudoir, 1837 *Coptopterella* Basilewsky, 1961 *Coptopterina* Basilewsky, 1956 *Crassagena* Baehr, 2006 *Cylindropectus* Lorenz, 1998 *Dolichoctis* Schmidt-Goebel, 1846 *Dontolobus* Basilewsky, 1970 *Drymatus* Motschulsky, 1862 *Eucheila* Dejean, 1829 *Eurycoleus* Chaudoir, 1848 *Eurydera* Laporte, 1831 *Formosiella* Jedlicka, 1951 *Gidda* Andrewes, 1920 *Glyphodactyla* Chaudoir, 1837 *Holcoderus* Chaudoir, 1870 *Horniulus* Jedlicka, 1932 *Labocephalus* Chaudoir, 1848 *Lelis* Chaudoir, 1870 *Lioptera* Chaudoir, 1870 *Lobodontidius* Basilewsky, 1970 *Lobodontulus* Basilewsky, 1970 *Lobodontus* Chaudoir, 1842 *Madecassina* Jeannel, 1949 *Mascarenhia* Alluaud, 1933 *Menarus* Andrewes, 1939 *Metascopus* Basilewsky, 1970 *Minuphloeus* Darlington, 1968 *Minuthodes* Andrewes, 1941 *Miscelus* Klug, 1834 *Mochtherus* Schmidt-Goebel, 1846 *Mormolyce* Hagenbach, 1825 *Mormolycina* Jeannel, 1949 *Neocoptodera* Jeannel, 1949 *Nycteis* Laporte, 1835 *Oreodicastes* Maindron, 1905 *Oxyodontus* Chaudoir, 1870 *Paradolichoctis* Baehr, 2006 *Pareurydera* Jeannel, 1949 *Pectinitarsus* Fairmaire, 1881 *Pericalus* W.S.MacLeay, 1825 *Philophlaeus* Chaudoir, 1844 *Phloeoxena* Chaudoir, 1870 *Pristacrus* Chaudoir, 1870 *Pseudoplatiella* Baehr, 2012 *Serrimargo* Chaudoir, 1870 *Sfitakantha* Andrewes, 1919 *Sinurus* Chaudoir, 1870 *Stenognathus* Chaudoir, 1843 *Stenotelus* Bouchard, 1903 *Stilboma* Andrewes, 1933 *Stricklandiana* Bousquet, 2002 *Tantillus* Chaudoir, 1870 *Thyreochaetus* Basilewsky, 1959 *Thyreopterus* Dejean, 1831 *Thysanotus* Chaudoir, 1848 *Trichocoptodera* Louwerens, 1958 *Xanthos* Kirschenhofer, 2003 *Xenitenopsis* Basilewsky, 1956 *Xenitenus* Péringuey, 1896 Subtribe Physoderina Chaudoir, 1877 *Allocota* Motschulsky, 1860 *Anchista* Nietner, 1856 *Dasiosoma* Britton, 1937 *Diamella* Shi & Liang, 2013 *Endynomena* Chaudoir, 1873 *Lachnoderma* W.J.MacLeay, 1873 *Metallanchista* Shi & Liang, 2013 *Orionella* Jedlicka, 1964 *Paraphaea* Bates, 1873 *Physodera* Eschscholtz, 1829 Subtribe Pseudotrechina Basilewsky, 1984 *Pseudotrechus* Rosenhauer, 1856 Subtribe Singilina Jeannel, 1949 *Singilis* Rambur, 1837 Subtribe Somotrichina Mateu, 1963 *Oecornis* Britton, 1940 *Paulianites* Jeannel, 1949 *Pephrica* Alluaud, 1936 *Somotrichus* Seidlitz, 1887 Subtribe Sugimotoina Habu, 1975 *Sugimotoa* Habu, 1975 Subtribe Trichina Basilewsky, 1984 *Trichis* Klug, 1832 *Cymindoides* Motschulsky, 1856 *Lebina* Germar, 1813 *Miolebidia* Zhang; Liu & Shangguan, 1989 *Protoscalidion* L.Schaufuss, 1888 Tribe Odacanthini Laporte, 1834 Subtribe Actenonycina Bates, 1871 *Actenonyx* White, 1846 Subtribe Homethina Liebherr, 2016 *Aeolodermus* Andrewes, 1929 *Cyphocoleus* Chaudoir, 1877 *Diplacanthogaster* Liebke, 1932 *Homethes* Newman, 1842 *Quammenis* Erwin, 2000 *Stenocheila* Laporte, 1832 Subtribe Odacanthina Laporte, 1834 *Anasis* Laporte, 1867 *Andrewesia* Liebke, 1938 *Arame* Andrewes, 1919 *Archicolliuris* Liebke, 1931 *Asios* Liebke, 1933 *Aulacolius* Sloane, 1923 *Basistichus* Sloane, 1917 *Clarencia* Sloane, 1917 *Colliuris* DeGeer, 1774 *Crassacantha* Baehr, 1995 *Cryptocolliuris* Basilewsky, 1955 *Deipyrodes* Bousquet, 2002 *Dicraspeda* Chaudoir, 1863 *Dobodura* Darlington, 1968 *Erectocolliuris* Liebke, 1931 *Eucolliuris* Liebke, 1931 *Eudalia* Laporte, 1867 *Gestroania* Liebke, 1938 *Giachinoana* Baehr, 2003 *Lachnothorax* Motschulsky, 1862 *Lasiocera* Dejean, 1831 *Mimocolliuris* Liebke, 1933 *Myrmecodemus* Sloane, 1923 *Neoeudalia* Baehr, 2005 *Odacantha* Paykull, 1798 *Ophionea* Klug, 1821 *Polydamasium* Liebke, 1938 *Porocara* Sloane, 1917 *Protocolliuris* Liebke, 1931 *Renneria* Baehr, 1999 *Smeringocera* Chaudoir, 1863 *Stenidia* Brullé, 1834 *Tricharnhemia* Baehr, 2009 Subtribe Pentagonicina Bates, 1873 *Parascopodes* Darlington, 1968 *Pentagonica* Schmidt-Goebel, 1846 *Scopodes* Erichson, 1842 Tribe Perigonini G.Horn, 1881 *Diploharpus* Chaudoir, 1850 *Mizotrechus* Bates, 1872 *Perigona* Laporte, 1835 *Ripogenites* Basilewsky, 1954

### Subfamily Licininae Bonelli, 1810

Tribe Chaetogenyini Emden, 1958 *Camptotoma* Reiche, 1843 Tribe Chlaeniini Brullé, 1834 Subtribe Callistina Laporte, 1834 *Callistomimus* Chaudoir, 1872 *Callistus* Bonelli, 1810 Subtribe Chlaeniina Brullé, 1834 *Actodus* Alluaud, 1915 *Chlaenius* Bonelli, 1810 *Eccoptomenus* Chaudoir, 1850 *Ectenognathus* Murray, 1858 *Globulipalpus* Sciaky & Facchini, 2019 *Harpaglossus* Motschulsky, 1858 *Hololeius* LaFerté-Sénectère, 1851 *Holosoma* Semenov, 1889 *Mirachlaenius* Facchini, 2011 *Parachlaenius* Kolbe, 1894 *Perissostomus* Alluaud, 1930 *Procletodema* Péringuey, 1898 *Procletus* Péringuey, 1896 *Rhopalomelus* Boheman, 1848 *Sphodroschema* Alluaud, 1930 *Stenoodes* Basilewsky, 1953 *Straneomelus* Sciaky & Facchini, 2019 *Stuhlmannium* Kolbe, 1894 *Viridagonum* Lassalle, 2015 *Rhopalochlaenius* Zhang; Sun & Zhang, 1994 Tribe Licinini Bonelli, 1810 Subtribe Dicaelina Laporte, 1834 *Dicaelus* Bonelli, 1813 *Diplocheila* Brullé, 1835 Subtribe Dicrochilina Ball, 1992 *Dicrochile* Guérin-Méneville, 1846 Subtribe Lestignathina Ball, 1992 *Atrotus* Péringuey, 1896 *Dilonchus* Andrewes, 1936 *Hormacrus* Sloane, 1898 *Lacordairia* Laporte, 1867 *Lestignathus* Erichson, 1842 *Microferonia* Blackburn, 1890 *Microzargus* Sciaky & Facchini, 1997 *Platylytron* W.J.MacLeay, 1873 *Siagonyx* W.J.MacLeay, 1871 *Zargus* Wollaston, 1854 Subtribe Licinina Bonelli, 1810 *Badister* Clairville, 1806 *Colpostoma* Semenov, 1889 *Derostichus* Motschulsky, 1860 *Eurygnathus* Wollaston, 1854 *Eutogeneius* Solier, 1849 *Licinus* Latreille, 1802 *Omestes* Andrewes, 1933 *Physolaesthus* Chaudoir, 1850 Tribe Oodini LaFerté-Sénectère, 1851 Subtribe Dercylina Sloane, 1923 *Dercylus* Laporte, 1832 Subtribe Geobaenina Péringuey, 1896 *Geobaenus* Dejean, 1829 Subtribe Melanchitonina Jeannel, 1948 *Dicaelindus* W.S.MacLeay, 1825 *Melanchiton* Andrewes, 1940 *Melanchrous* Andrewes, 1940 Subtribe Oodina LaFerté-Sénectère, 1851 *Acanthoodes* Basilewsky, 1953 *Acutosternus* Lecordier & Girard, 1988 *Adelopomorpha* Heller, 1916 *Anatrichis* LeConte, 1853 *Bamaroodes* Gueorguiev, 2014 *Brachyodes* Jeannel, 1949 *Chaetocrepis* Chaudoir, 1857 *Coptocarpus* Chaudoir, 1857 *Dercylinus* Chaudoir, 1883 *Evolenes* LeConte, 1853 *Holcocoleus* Chaudoir, 1883 *Hoplolenus* LaFerté-Sénectère, 1851 *Lobatodes* Basilewsky, 1956 *Lonchosternus* LaFerté-Sénectère, 1851 *Macroprotus* Chaudoir, 1878 *Megaloodes* Lesne, 1896 *Microodes* Jeannel, 1949 *Miltodes* Andrewes, 1922 *Nanodiodes* Bousquet, 1996 *Neoodes* Basilewsky, 1953 *Nothoodes* Gueorguiev & Liang, 2020 *Oodes* Bonelli, 1810 *Oodinus* Motschulsky, 1865 *Orthocerodus* Basilewsky, 1946 *Polychaetus* Chaudoir, 1882 *Prionognathus* LaFerté-Sénectère, 1851 *Protopidius* Basilewsky, 1949 *Pseudoodes* Gueorguiev & Liang, 2020 *Pseudosphaerodes* Jeannel, 1949 *Simous* Chaudoir, 1882 *Sphaerodinus* Jeannel, 1949 *Sphoerodes* Chaudoir, 1883 *Stenocrepis* Chaudoir, 1857 *Sundaoodes* Gueorguiev & Liang, 2020 *Systolocranius* Chaudoir, 1857 *Thaioodes* Gueorguiev, 2014 *Thryptocerus* Chaudoir, 1878 *Trichopalpoodes* B.Gueorguiev & J.Schmidt, 2016

### Subfamily Loricerinae Bonelli, 1810

*Loricera* Latreille, 1802

### Subfamily Melaeninae Csiki, 1933

*Cymbionotum* Baudi di Selve, 1864 *Melaenus* Dejean, 1831

### Subfamily Migadopinae Chaudoir, 1861

Tribe Amarotypini Erwin, 1985 *Amarotypus* Bates, 1872 *Migadopiella* Baehr, 2009 Tribe Migadopini Chaudoir, 1861 *Antarctonomus* Chaudoir, 1861 *Aquilex* Moret, 1989 *Calathosoma* Jeannel, 1938 *Calyptogonia* Sloane, 1920 *Decogmus* Sloane, 1915 *Dendromigadops* Baehr, 2013 *Lissopterus* G.R.Waterhouse, 1843 *Loxomerus* Chaudoir, 1842 *Migadopidius* Jeannel, 1938 *Migadops* G.R.Waterhouse, 1842 *Monolobus* Solier, 1849 *Nebriosoma* Laporte, 1867 *Pseudomigadops* Jeannel, 1938 *Rhytidognathus* Chaudoir, 1861 *Stichonotus* Sloane, 1910 *Taenarthrus* Broun, 1914

### Subfamily Nebriinae Laporte, 1834

Tribe Cicindini Csiki, 1927 *Archaeocindis* Kavanaugh & Erwin, 1991 *Cicindis* Bruch, 1908 Tribe Nebriini Laporte, 1834 *Leistus* Frölich, 1799 *Nebria* Latreille, 1802 *Nippononebria* Ueno, 1955 *Archaeonebria* Kavanaugh & J.Schmidt, 2019 *Ledouxnebria* Deuve, 1998 Tribe Notiokasiini Kavanaugh & Nègre, 1983 *Notiokasis* Kavanaugh & Nègre, 1983 Tribe Notiophilini Motschulsky, 1850 *Notiophilus* Duméril, 1805 Tribe Opisthiini Dupuis, 1912 *Opisthius* Kirby, 1837 *Paropisthius* Casey, 1920 Tribe Pelophilini Kavanaugh, 1996 *Pelophila* Dejean, 1821

### Subfamily Nototylinae Bänninger, 1927

*Nototylus* Gemminger & Harold, 1868

### Subfamily Omophroninae Bonelli, 1810

*Omophron* Latreille, 1802

### Subfamily Orthogoniinae Schaum, 1857

Tribe Amorphomerini Sloane, 1923 *Amorphomerus* Sloane, 1923 Tribe Idiomorphini Bates, 1891 Subtribe Idiomorphina Bates, 1891 *Idiomorphus* Chaudoir, 1846 *Rathymus* Dejean, 1831 *Strigia* Brullé, 1835 Subtribe Perochnoristhina Basilewsky, 1973 *Perochnoristhus* Basilewsky, 1973 Tribe Orthogoniini Schaum, 1857 Subtribe Glyptina G.Horn, 1881 *Glyptus* Brullé, 1837 *Neoglyptus* Basilewsky, 1953 Subtribe Orthogoniina Schaum, 1857 *Actenoncus* Chaudoir, 1872 *Anoncopeucus* Chaudoir, 1872 *Hexachaetus* Chaudoir, 1872 *Neoorthogonius* Tian & Deuve, 2006 *Nepalorthogonius* Habu, 1979 *Orthogonius* W.S.MacLeay, 1825

### Subfamily Panagaeinae Bonelli, 1810

Tribe Brachygnathini Basilewsky, 1946 *Brachygnathus* Perty, 1830 Tribe Panagaeini Bonelli, 1810 Subtribe Bascanina Basilewsky, 1953 *Bascanus* Péringuey, 1896 Subtribe Panagaeina Bonelli, 1810 *Adischissus* Fedorenko, 2015 *Calathocosmus* Emden, 1928 *Cintaroa* Kasahara, 1989 *Cintaromorpha* Häckel & Anichtchenko, 2016 *Coptia* Brullé, 1835 *Craspedophorus* Hope, 1838 *Dischissus* Bates, 1873 *Epigraphodes* Basilewsky, 1967 *Epigraphus* Chaudoir, 1869 *Euschizomerus* Chaudoir, 1850 *Geobius* Dejean, 1831 *Micrixys* LeConte, 1854 *Microschemus* Andrewes, 1940 *Panagaeus* Latreille, 1802 *Paregraphus* Basilewsky, 1967 *Peronomerus* Schaum, 1854 *Psecadius* Alluaud, 1911 *Tefflus* Leach in Samouelle, 1819 *Tinoderus* Chaudoir, 1879 *Tinognathus* Chaudoir, 1879 *Trichisia* Motschulsky, 1865 Tribe Peleciini Chaudoir, 1880 Subtribe Agonicina Sloane, 1920 *Agonica* Sloane, 1920 *Pseudagonica* B.Moore, 1960 Subtribe Peleciina Chaudoir, 1880 *Ardistomopsis* Straneo & Ball, 1989 *Disphericus* G.R.Waterhouse, 1842 *Dyschiridium* Chaudoir, 1861 *Eripus* Dejean, 1829 *Pelecium* Kirby, 1819 *Stricteripus* Straneo & Ball, 1989

### Subfamily Patrobinae Kirby, 1837

Tribe Lissopogonini Zamotajlov, 2000 *Lissopogonus* Andrewes, 1923 Tribe Patrobini Kirby, 1837 Subtribe Deltomerina Chaudoir, 1872 *Deltomerus* Motschulsky, 1850 Subtribe Deltomerodina Zamotajlov, 2002 *Deltomerodes* Deuve, 1992 Subtribe Patrobina Kirby, 1837 *Apatrobus* Habu & Baba, 1960 *Apenetretus* Kurnakov, 1960 *Archipatrobus* Zamotajlov, 1992 *Caelopenetretus* Zamotajlov & Ito, 2000 *Chaetapatrobus* Lafer, 1996 *Chinapenetretus* Kurnakov, 1963 *Dimorphopatrobus* Casale & Sciaky, 1994 *Diplous* Motschulsky, 1850 *Grandipenetretus* Zamotajlov & Sciaky, 1999 *Himalopenetretus* Zamotajlov, 2002 *Indopatrobus* Zamotajlov & Wrase, 2006 *Ledouxius* Zamotajlov, 1992 *Minipenetretus* Zamotajlov, 2002 *Minypatrobus* Ueno, 1955 *Naxipenetretus* Zamotajlov, 1999 *Parapatrobus* Zamotajlov, 1992 *Parapenetretus* Kurnakov, 1960 *Patanitretus* Zamotajlov, 2002 *Patrobus* Dejean, 1821 *Penetretus* Motschulsky, 1865 *Platidiolus* Chaudoir, 1878 *Platypatrobus* Darlington, 1938 *Prodiplous* Zamotajlov & Sciaky, 2006 *Qiangopatrobus* Zamotajlov, 2002 *Quasipenetretus* Zamotajlov, 2002 *Tibetopenetretus* Zamotajlov & Sciaky, 1999

### Subfamily Paussinae Latreille, 1806

Tribe Metriini LeConte, 1853 Subtribe †Kryzhanovskianina Deuve, 2019 *Kryzhanovskiana* Kataev & Kirejtshuk, 2019 *Metrius* Eschscholtz, 1829 *Sinometrius* Wrase & J.Schmidt, 2006 Tribe Ozaenini Hope, 1838 Subtribe Microzaenina Deuve, 2019 *Microzaena* Fairmaire, 1901 Subtribe Mystropomina G.Horn, 1881 *Mystropomus* Chaudoir, 1848 Subtribe Ozaenina Hope, 1838 *Crepidozaena* Deuve, 2001 *Entomoantyx* Ball & McCleve, 1990 *Filicerozaena* Deuve, 2001 *Gibbozaena* Deuve, 2001 *Goniotropis* G.R.Gray, 1831 *Inflatozaena* Deuve, 2001 *Mimozaena* Deuve, 2001 *Ozaena* Olivier, 1812 *Pachyteles* Perty, 1830 *Physea* Brullé, 1835 *Physeomorpha* Ogueta, 1963 *Platycerozaena* Bänninger, 1927 *Proozaena* Deuve, 2001 *Serratozaena* Deuve, 2001 *Tachypeles* Deuve, 2001 *Tropopsis* Solier, 1849 Subtribe Pseudozaenina Sloane, 1905 *Anentmetus* Andrewes, 1924 *Dhanya* Andrewes, 1919 *Eustra* Schmidt-Goebel, 1846 *Itamus* Schmidt-Goebel, 1846 *Pseudozaena* Laporte, 1834 *Sphaerostylus* Chaudoir, 1848 Tribe Paussini Latreille, 1806 Subtribe Carabidomemnina Wasmann, 1928 *Carabidomemnus* Kolbe, 1924 *Eohomopterus* Wasmann, 1919 Subtribe Cerapterina Billberg, 1820 *Arthropterus* W.S.MacLeay, 1838 *Cerapterus* Swederus, 1788 *Megalopaussus* Lea, 1906 *Mesarthropterus* Wasmann, 1926 Subtribe Heteropaussina Janssens, 1950 *Heteropaussus* J.Thomson, 1860 Subtribe Homopterina Wasmann, 1920 *Homopterus* Westwood, 1841 Subtribe Paussina Latreille, 1806 *Allorhopalus* Nagel, 2017 *Ceratoderus* Westwood, 1841 *Euplatyrhopalus* Desneux, 1905 *Lebioderus* Westwood, 1838 *Leleupaussus* Luna de Carvalho, 1962 *Melanospilus* Westwood, 1847 *Paussomorphus* Raffray, 1885 *Paussus* Linnaeus, 1775 *Platyrhopalides* Wasmann, 1918 *Platyrhopalopsis* Desneux, 1905 *Platyrhopalus* Westwood, 1833 *Pterorhopalus* Maruyama, 2011 Subtribe Pentaplatarthrina Jeannel, 1946 *Hexaplatarthrus* Jeannel, 1955 *Pentaplatarthrus* Westwood, 1833 Subtribe †Arthropteritina Luna de Carvalho, 1961 *Arthropterites* Wasmann, 1926 *Cerapterites* Wasmann, 1926 *Protocerapterus* Wasmann, 1926 Subtribe †Eopaussina Luna de Carvalho, 1951 *Eopaussus* Wasmann, 1926 Tribe Protopaussini Gestro, 1892 *Protopaussus* Gestro, 1892

### Subfamily Platyninae Bonelli, 1810

Tribe Omphreini Ganglbauer, 1891 *Omphreus* Dejean, 1828 Tribe Platynini Bonelli, 1810 Subtribe Enoicina Basilewsky, 1985 *Abacetodes* Straneo, 1939 *Enoicus* Péringuey, 1896 Subtribe Platynina Bonelli, 1810 *Achaetocephala* Habu, 1975 *Achaetoprothorax* Habu, 1978 *Aepsera* Chaudoir, 1874 *Agelaea* Gené, 1839 *Agonidium* Jeannel, 1948 *Agonobembix* Jeannel, 1948 *Agonoriascus* Basilewsky, 1985 *Agonorites* Jeannel, 1951 *Agonum* Bonelli, 1810 *Altagonum* Darlington, 1952 *Anchomenus* Bonelli, 1810 *Andinocolpodes* Perrault, 1991 *Andrewesius* Andrewes, 1939 *Aparupa* Andrewes, 1930 *Archagonum* Basilewsky, 1953 *Archicolpodes* J.Schmidt, 2001 *Arhytinus* Bates, 1889 *Atranodes* Jedlicka, 1953 *Atranus* LeConte, 1847 *Austroglyptolenus* Roig-Juñent, 2003 *Beckeria* Jedlicka, 1931 *Blackburnia* Sharp, 1878 *Bothrocolpodes* Basilewsky, 1985 *Bruskespar* Morvan, 1998 *Bruskmoal* Morvan, 1998 *Bryanites* Valentine, 1987 *Callidagonum* Lorenz, 1998 *Cardiomera* Bassi, 1834 *Catacolpodes* Basilewsky, 1985 *Celaenagonum* Habu, 1978 *Chaetagonum* Burgeon, 1933 *Chaetosaurus* J.Schmidt, 2001 *Cinctagonum* Baehr, 2012 *Cistelagonum* Baehr, 2012 *Collagonum* Baehr, 1995 *Colpodes* W.S.MacLeay, 1825 *Colpoides* Jedlicka, 1931 *Colpomimus* Basilewsky, 1985 *Colposphodrus* Jedlicka, 1953 *Ctenognathus* Fairmaire, 1843 *Cymenopterus* Jeannel, 1948 *Cyrtopilus* Basilewsky, 1985 *Dalatagonum* Fedorenko, 2011 *Deliaesianum* Morvan, 1999 *Deltocolpodes* Morvan, 1992 *Dendragonum* Jeannel, 1948 *Diacanthostylus* Habu, 1978 *Dicranoncoides* Habu, 1978 *Dicranoncus* Chaudoir, 1850 *Dinocolpodes* J.Schmidt, 2001 *Dirotus* W.S.MacLeay, 1825 *Dister* Morvan, 2006 *Dolichocolpodes* Basilewsky, 1985 *Dyscolus* Dejean, 1831 *Elliptoleus* Bates, 1882 *Epicolpodes* Basilewsky, 1985 *Eucolpodes* Jeannel, 1948 *Euleptus* Klug, 1833 *Euplynes* Schmidt-Goebel, 1846 *Feroniascus* Jeannel, 1951 *Fortagonum* Darlington, 1952 *Galiciotyphlotes* Assmann, 1999 *Gastragonum* Darlington, 1952 *Glaucagonum* Habu, 1978 *Glyptolenoides* Perrault, 1991 *Glyptolenus* Bates, 1878 *Gyrochaetostylus* Habu, 1978 *Habragonum* Ueno, 1964 *Hannaphota* Landin, 1955 *Haplocolpodes* Jeannel, 1951 *Haplopeza* Boheman, 1848 *Helluocolpodes* Liebherr, 2005 *Hemiplatynus* Casey, 1920 *Henvelik* Morvan, 1999 *Herculagonum* Baehr, 2002 *Hikosanoagonum* Habu, 1954 *Idiagonum* Darlington, 1952 *Idiastes* Andrewes, 1931 *Idiocolpodes* Basilewsky, 1985 *Incagonum* Liebherr, 1994 *Iridagonum* Darlington, 1952 *Ischnagonum* Kasahara & Satô, 1997 *Jocqueius* Basilewsky, 1988 *Jujiroa* Ueno, 1952 *Kalchdigor* Morvan, 1999 *Kar* Morvan, 1998 *Karnes* Morvan, 2010 *Kaszabellus* Jedlicka, 1954 *Kiwiplatynus* Larochelle & Larivière, 2021 *Klapperichella* Jedlicka, 1956 *Kuceraianum* Morvan, 2002 *Kupeplatynus* Larochelle & Larivière, 2021 *Laevagonum* Darlington, 1952 *Lassalleianum* Morvan, 1999 *Lepcha* Andrewes, 1930 *Leptagonum* Kolbe, 1897 *Leptocolpodes* Basilewsky, 1985 *Letouzeya* Bruneau de Miré, 1982 *Liagonum* Jeannel, 1948 *Liamegalonychus* Basilewsky, 1963 *Liocolpodes* Basilewsky, 1985 *Lissagonum* Habu, 1978 *Lithagonum* Darlington, 1952 *Lobocolpodes* Basilewsky, 1985 *Lorostema* Motschulsky, 1865 *Loxocrepis* Eschscholtz, 1829 *Lucicolpodes* J.Schmidt, 2000 *Maculagonum* Darlington, 1952 *Maoriplatynus* Larochelle & Larivière, 2021 *Meleagros* Kirschenhofer, 1999 *Mesocolpodes* Basilewsky, 1985 *Metacolpodes* Jeannel, 1948 *Mexisphodrus* Barr, 1965 *Montagonum* Darlington, 1952 *Mooreagonum* Baehr, 2016 *Morimotoidius* Habu, 1954 *Nebriagonum* Darlington, 1952 *Negreum* Habu, 1958 *Neobatenus* Jeannel, 1948 *Neocolpodes* Jeannel, 1948 *Neodendragonum* Basilewsky, 1953 *Neomegalonychus* Jeannel, 1948 *Nesiocolpodes* Jeannel, 1948 *Nipponagonum* Habu, 1978 *Notagonum* Darlington, 1952 *Notocolpodes* Basilewsky, 1985 *Notoplatynus* B.Moore, 1985 *Nymphagonum* Habu, 1978 *Olisthopus* Dejean, 1828 *Oncostylus* Habu, 1978 *Onotokiba* Alluaud, 1927 *Onycholabis* Bates, 1873 *Onypterygia* Dejean, 1831 *Orophicus* Alluaud, 1925 *Orthotrichus* Peyron, 1856 *Oxygonium* Basilewsky, 1951 *Oxypselaphus* Chaudoir, 1843 *Pachybatenus* Basilewsky, 1973 *Pachyferonia* Jeannel, 1951 *Paracolpodes* Basilewsky, 1985 *Paraliagonum* Basilewsky, 1957 *Paramegalonychus* Basilewsky, 1953 *Paranchodemus* Habu, 1978 *Paranchus* Lindroth, 1974 *Paraplatynus* Baehr, 2016 *Pawgammm* Morvan, 1999 *Platyagonum* Habu, 1978 *Platynus* Bonelli, 1810 *Plaumannium* Liebke, 1939 *Plicagonum* Darlington, 1952 *Ponapagonum* Darlington, 1970 *Potamagonum* Darlington, 1952 *Praepristus* Kirschenhofer, 1999 *Promecoptera* Dejean, 1831 *Promegalonychus* Basilewsky, 1953 *Prophenorites* Basilewsky, 1985 *Prosphodrus* Britton, 1959 *Protocolpodes* Basilewsky, 1985 *Pseudanchomenus* Tarnier, 1860 *Pseudobatenus* Basilewsky, 1951 *Pseudomegalonychus* Basilewsky, 1950 *Rhadine* LeConte, 1846 *Rupa* Jedlicka, 1935 *Scotagonum* Habu, 1978 *Sericoda* Kirby, 1837 *Shibataia* Habu, 1978 *Sinocolpodes* J.Schmidt, 2001 *Skoeda* Morvan, 1995 *Skorlagad* Morvan, 1999 *Skouedhirraad* Morvan, 1999 *Sophroferonia* Alluaud, 1933 *Speagonum* B.Moore, 1977 *Speocolpodes* Barr, 1974 *Speokokosia* Alluaud, 1932 *Sperkanhir* Morvan, 2010 *Stenocnemus* Mannerheim, 1837 *Straneoa* Basilewsky, 1953 *Syletor* Tschitscherine, 1899 *Takasagoagonum* Habu, 1977 *Tanystoma* Motschulsky, 1845 *Tarsagonum* Darlington, 1952 *Tetraleucus* Casey, 1920 *Tostkar* Morvan, 1998 *Trogloagonum* Casale, 1982 *Tuiplatynus* Larochelle & Larivière, 2021 *Vitagonum* B.Moore, 1999 *Vulcanophilus* Heller, 1898 *Xestagonum* Habu, 1978 *Praeanchodemus* J.Schmidt; Göpel & Will, 2017 Tribe Sphodrini Laporte, 1834 Subtribe Atranopsina Baehr, 1982 *Amaroschema* Jeannel, 1943 *Amphimasoreus* Piochard de la Brûlerie, 1875 *Atranopsis* Baehr, 1982 *Broter* Andrewes, 1923 *Gomerina* Bolivar y Pieltain, 1940 *Paraeutrichopus* Mateu, 1954 *Platyderus* Stephens, 1827 *Pseudomyas* Uyttenboogaart, 1929 *Pseudoplatyderus* Bolivar y Pieltain, 1940 Subtribe Calathina Laporte, 1834 *Calathus* Bonelli, 1810 *Thermoscelis* Putzeys, 1873 Subtribe Dolichina Brullé, 1834 *Acalathus* Semenov, 1889 *Anchomenidius* Heyden, 1880 *Casaleius* Sciaky & Wrase, 1998 *Dolichus* Bonelli, 1810 *Doliodactyla* Sciaky & Wrase, 1998 *Morphodactyla* Semenov, 1889 *Xestopus* Andrewes, 1937 Subtribe Pristosiina Lindroth, 1956 *Pristosia* Motschulsky, 1865 Subtribe Sphodrina Laporte, 1834 *Calathidius* Putzeys, 1873 *Cephalosdrophus* Lassalle & Marcilhac, 1999 *Eosphodrus* Casale, 1988 *Eremosphodrus* Semenov, 1909 *Himalosphodrus* Casale, 1988 *Hystricosphodrus* Casale & Giachino, 2004 *Ifridytes* Deuve & Queinnec, 1994 *Laemostenus* Bonelli, 1810 *Licinopsis* Bedel, 1899 *Miquihuana* Barr, 1982 *Pseudotaphoxenus* L.Schaufuss, 1865 *Reflexisphodrus* Casale, 1988 *Sphodropsis* Seidlitz, 1887 *Sphodrus* Clairville, 1806 *Stenolepta* Semenov, 1889 *Taphoxenus* Motschulsky, 1850 Subtribe Synuchina Lindroth, 1956 *Nipponosynuchus* Morita, 1998 *Parabroscus* Lindroth, 1956 *Synuchus* Gyllenhal, 1810 *Trephionus* Bates, 1883

### Subfamily Promecognathinae LeConte, 1853

Tribe Axinidiini Basilewsky, 1963 *Axinidium* Sturm, 1843 *Holaxinidium* Basilewsky, 1963 *Metaxinidium* Basilewsky, 1963 *Paraxinidium* Basilewsky, 1963 Tribe Dalyatini Mateu, 2002 *Dalyat* Mateu, 2002 Tribe Promecognathini LeConte, 1853 *Promecognathus* Chaudoir, 1846 Tribe †Palaeoaxinidiini McKay, 1991 *Palaeoaxinidium* McKay, 1991

### Subfamily Pseudomorphinae Hope, 1838

*Adelotopus* Hope, 1836 *Cainogenion* Notman, 1925 *Cryptocephalomorpha* Ritsema, 1875 *Guyanemorpha* Erwin, 2013 *Manumorpha* Erwin & Geraci, 2008 *Notopseudomorpha* Baehr, 1997 *Paussotropus* C.O.Waterhouse, 1877 *Pseudomorpha* Kirby, 1823 *Samiriamorpha* Erwin & Geraci, 2008 *Sphallomorpha* Westwood, 1840 *Tuxtlamorpha* Erwin & Geraci, 2008 *Yasunimorpha* Erwin & Geraci, 2008

### Subfamily Psydrinae LeConte, 1853

Tribe Gehringiini Darlington, 1933 Subtribe Gehringiina Darlington, 1933 *Gehringia* Darlington, 1933 Subtribe Helenaeina Deuve, 2007 *Afrogehringia* Baehr; Schüle & Lorenz, 2009 *Helenaea* Schatzmayr & Koch, 1934 Tribe Moriomorphini Sloane, 1890 Subtribe Amblytelina Blackburn, 1892 *Amblytelus* Erichson, 1842 *Dystrichothorax* Blackburn, 1892 *Epelyx* Blackburn, 1892 *Mecyclothorax* Sharp, 1903 *Meonis* Laporte, 1867 *Paratrichothorax* Baehr, 2004 *Pseudamblytelus* Baehr, 2004 *Raphetis* B.Moore, 1963 *Selenochilus* Chaudoir, 1878 *Trichamblytelus* Baehr, 2004 Subtribe Moriomorphina Sloane, 1890 *Celanida* Laporte, 1867 *Melisodera* Westwood, 1835 *Molopsida* White, 1846 *Moriodema* Laporte, 1867 *Moriomorpha* Laporte, 1867 *Neonomius* B.Moore, 1963 *Pterogmus* Sloane, 1920 *Rhaebolestes* Sloane, 1903 *Rossjoycea* Liebherr, 2011 *Sitaphe* B.Moore, 1963 *Spherita* Liebherr, 2020 *Tarastethus* Sharp, 1883 *Teraphis* Laporte, 1867 *Theprisa* B.Moore, 1963 *Trephisa* B.Moore, 1963 *Trichopsida* Larochelle & Larivière, 2013 Subtribe Tropopterina Sloane, 1898 *Pharetis* Liebherr, 2020 *Tropopterus* Solier, 1849 Tribe Psydrini LeConte, 1853 *Laccocenus* Sloane, 1890 *Nomius* Laporte, 1835 *Psydrus* LeConte, 1846

### Subfamily Pterostichinae Bonelli, 1810

Tribe Chaetodactylini Tschitscherine, 1903 *Chaetodactyla* Tschitscherine, 1897 Tribe Cnemalobini Germain, 1911 *Cnemalobus* Guérin-Méneville, 1838 Tribe Cratocerini Lacordaire, 1854 Subtribe Catapieseina Bates, 1882 *Catapiesis* Solier, 1835 *Homalomorpha* Brullé, 1837 Subtribe Cratocerina Lacordaire, 1854 *Brachidius* Chaudoir, 1852 *Cratocerus* Dejean, 1829 *Oxyglychus* Straneo, 1938 Subtribe Drimostomatina Chaudoir, 1873 *Abacaelostus* Straneo, 1952 *Andrewesinulus* Straneo, 1938 *Apsidocnemus* Alluaud, 1936 *Barylaus* Liebherr, 1985 *Caecocaelus* Straneo, 1949 *Caelostomus* W.S.MacLeay, 1825 *Camptogenys* Tschitscherine, 1899 *Capabatus* Csiki, 1930 *Crenulostrigus* Straneo, 1942 *Cyrtolaus* Bates, 1882 *Dactyleurys* Tschitscherine, 1899 *Dactylinius* Straneo, 1941 *Diachipteryx* Alluaud, 1925 *Diceromerus* Chaudoir, 1873 *Dromistomus* Jeannel, 1948 *Drymonaxus* Straneo, 1941 *Feostoma* Straneo, 1941 *Hemitelestus* Alluaud, 1895 *Hoplizomenus* Chaudoir, 1873 *Leleuporites* Straneo, 1960 *Madapelmus* Dajoz, 1985 *Monodryxus* Straneo, 1942 *Pachycaecus* Straneo, 1971 *Pachyroxochus* Straneo, 1942 *Platyxythrius* Lorenz, 1998 *Stegazopteryx* Will, 2004 *Stomonaxellus* Tschitscherine, 1901 *Strigomerodes* Straneo, 1939 *Strigomerus* Chaudoir, 1873 *Trichillinus* Straneo, 1938 Tribe Microcheilini Jeannel, 1948 *Microcheila* Brullé, 1835 Tribe Morionini Brullé, 1837 *Buderes* Murray, 1857 *Hyperectenus* Alluaud, 1935 *Hyperion* Laporte, 1834 *Megamorio* Chaudoir, 1880 *Morion* Latreille, 1810 *Morionidius* Chaudoir, 1880 *Moriosomus* Motschulsky, 1855 *Platynodes* Westwood, 1847 *Stereostoma* Murray, 1857 Tribe Pterostichini Bonelli, 1810 Subtribe Abacetina Chaudoir, 1873 *Abacaecus* Allegro & Giachino, 2020 *Abacetus* Dejean, 1828 *Aristopus* LaFerté-Sénectère, 1853 *Celioinkosa* Straneo, 1951 *Chlaeminus* Motschulsky, 1865 *Colpodichius* Straneo, 1952 *Cosmodiscus* Sloane, 1907 *Cyrtomoscelis* Chaudoir, 1874 *Distrigidius* Jeannel, 1948 *Ecnomolaus* Bates, 1892 *Haptoderidius* Straneo, 1975 *Haptoderodes* Straneo, 1986 *Holconotus* Schmidt-Goebel, 1846 *Inkosa* Péringuey, 1926 *Mateuellus* Deuve, 1990 *Metabacetus* Bates, 1892 *Metaxellus* Straneo, 1960 *Metaxys* Chaudoir, 1857 *Novillidius* Straneo, 1941 *Oodinkosa* Straneo, 1939 *Ophonichius* Straneo, 1942 *Pediomorphus* Chaudoir, 1878 *Pioprosopus* Tschitscherine, 1899 *Pollicobius* Vinson, 1939 *Prostalomus* Basilewsky, 1950 *Pseudabacetus* Burgeon, 1935 *Pterostillichus* Straneo, 1949 *Rhagadillius* Straneo, 1951 *Tiferonia* Darlington, 1962 *Trachelocyphoides* Straneo, 1942 *Trachelocyphus* Tschitscherine, 1900 Subtribe Euchroina Chaudoir, 1874 *Abacillius* Straneo, 1949 *Abacillodes* Straneo, 1988 *Abaris* Dejean, 1831 *Apsaustodon* Tschitscherine, 1901 *Argutoridius* Chaudoir, 1876 *Blennidus* Motschulsky, 1866 *Blenniventer* Straneo, 1986 *Bothynoproctus* Tschitscherine, 1900 *Cedrorum* Borges & A.Serrano, 1993 *Cephalostichus* Straneo, 1977 *Cynthidia* Chaudoir, 1874 *Euchroa* Brullé, 1835 *Eumara* Tschitscherine, 1901 *Gastrogmus* Sloane, 1915 *Haplobothynus* Tschitscherine, 1901 *Litarthrum* Sloane, 1915 *Lobobrachus* Sharp, 1885 *Marsyas* Putzeys, 1845 *Meropalpus* Tschitscherine, 1900 *Microcephalus* Dejean, 1828 *Neotalus* Will, 2002 *Oribazus* Chaudoir, 1874 *Orthomus* Chaudoir, 1838 *Pachythecus* Chaudoir, 1874 *Paniestichus* Will, 2011 *Parorthomus* B.Gueorguiev; Wrase & Farkac, 2014 *Phaenaulax* Tschitscherine, 1898 *Prosopogmus* Chaudoir, 1865 *Pseudabarys* Chaudoir, 1874 *Setalimorphus* Sloane, 1895 *Setalis* Laporte, 1867 *Simodontus* Chaudoir, 1843 *Trirammatus* Chaudoir, 1835 Subtribe Loxandrina Erwin & Sims, 1984 *Cerabilia* Laporte, 1867 *Oxycrepis* Reiche, 1843 *Zeodera* Laporte, 1867 Subtribe Metiina Straneo, 1951 *Abropus* G.R.Waterhouse, 1842 *Antarctiola* Straneo, 1977 *Feroniola* Tschitscherine, 1900 *Kuschelinus* Straneo, 1963 *Metius* Curtis, 1839 Subtribe Pterostichina Bonelli, 1810 *Abacidus* LeConte, 1863 *Abacoleptus* Fauvel, 1903 *Abacomorphus* Chaudoir, 1878 *Abacophrastus* Will, 2011 *Abacops* Tschitscherine, 1902 *Abax* Bonelli, 1810 *Allotriopus* Bates, 1882 *Analoma* Darlington, 1971 *Apophylon* B.Gueorguiev & Sciaky, 2015 *Aristochroa* Tschitscherine, 1898 *Aristochroodes* Marcilhac, 1993 *Aulacopodus* Britton, 1940 *Basilewskya* Straneo, 1948 *Camptoscelis* Dejean, 1828 *Castelnaudia* Tschitscherine, 1891 *Catadromus* W.S.MacLeay, 1825 *Chaetauchenium* Tschitscherine, 1900 *Chalcochrous* Chaudoir, 1838 *Cophosomorpha* Tschitscherine, 1891 *Cratoferonia* Tschitscherine, 1902 *Cratogaster* Blanchard, 1843 *Cuneipectus* Sloane, 1907 *Cyclotrachelus* Chaudoir, 1838 *Cyrtoderus* Hope, 1842 *Darodilia* Laporte, 1867 *Delinius* Westwood, 1864 *Eucamptognathus* Chaudoir, 1837 *Eudromus* Klug, 1835 *Euryabax* Fauvel, 1903 *Euryaptus* Bates, 1892 *Eurypercus* Jeannel, 1948 *Eurystomis* Chaudoir, 1878 *Gastrellarius* Casey, 1918 *Gourlayia* Britton, 1964 *Harpostomus* Chaudoir, 1856 *Henrotiochoromus* Busulini, 1958 *Henrotius* Jeannel, 1953 *Holcaspis* Chaudoir, 1865 *Hybothecus* Chaudoir, 1874 *Leiolesticus* Roux; Lassalle & Dubault, 2016 *Lesticus* Dejean, 1828 *Licentius* Jedlicka, 1939 *Liopasa* Tschitscherine, 1901 *Lophoglossus* LeConte, 1853 *Loxogenius* Sloane, 1907 *Mecynognathus* W.J.MacLeay, 1873 *Megadromus* Motschulsky, 1866 *Molopidius* Jeannel, 1942 *Molopinus* Jeannel, 1948 *Molops* Bonelli, 1810 *Myas* Sturm, 1826 *Neoferonia* Britton, 1940 *Nesites* Andrewes, 1939 *Nirmala* Andrewes, 1930 *Notabax* B.Moore, 1976 *Notolestus* Sloane, 1894 *Notonomus* Chaudoir, 1865 *Nurus* Motschulsky, 1866 *Oberthueria* Vuillet, 1911 *Ogmophora* Tschitscherine, 1896 *Onawea* Johns, 2007 *Oscadytes* Lagar Mascaro, 1975 *Pachymorphus* Chaudoir, 1838 *Paranurus* Tschitscherine, 1901 *Pareuryaptus* Dubault; Lassalle & Roux, 2008 *Parhypates* Motschulsky, 1866 *Pedius* Motschulsky, 1850 *Percolaus* Bates, 1882 *Percus* Bonelli, 1810 *Peyrieraselus* Deuve, 1981 *Piesmus* LeConte, 1846 *Platycoelus* Blanchard, 1843 *Platysmodes* Fauvel, 1903 *Plocamostethus* Britton, 1940 *Poecilinus* Jeannel, 1948 *Poecilus* Bonelli, 1810 *Pseudoceneus* Tschitscherine, 1891 *Pterostichus* Bonelli, 1810 *Rhabdotus* Chaudoir, 1865 *Rhytiferonia* Darlington, 1962 *Rhytisternus* Chaudoir, 1865 *Sarticus* Motschulsky, 1866 *Secatophus* Laporte, 1867 *Setalidius* Chaudoir, 1878 *Speluncarius* Reitter, 1886 *Speomolops* Patrizi, 1955 *Sphodrosomus* Perroud, 1864 *Stenochoromus* L.Miller, 1866 *Stereocerus* Kirby, 1837 *Sthenocranion* Tschitscherine, 1899 *Stomis* Clairville, 1806 *Straneostichus* Sciaky, 1994 *Styracoderus* Chaudoir, 1874 *Tanythrix* Schaum, 1858 *Taphoxomimus* Straneo, 1975 *Tapinopterus* Schaum, 1858 *Teratotarsa* Tschitscherine, 1893 *Teropha* Laporte, 1867 *Trichosternus* Chaudoir, 1865 *Trigonaptus* Fedorenko, 2020 *Trigonotoma* Dejean, 1828 *Troglorites* Jeannel, 1919 *Tropidocerus* Chaudoir, 1878 *Typhlochoromus* Moczarski, 1913 *Wahlbergiana* Bousquet, 2002 *Xenion* Tschitscherine, 1902 *Zariquieya* Jeannel, 1924 *Zeopoecilus* Sharp, 1886 Tribe Zabrini Bonelli, 1810 Subtribe Amarina Zimmermann, 1832 *Amara* Bonelli, 1810 *Pseudamara* Lindroth, 1968 Subtribe Zabrina Bonelli, 1810 *Zabrus* Clairville, 1806

### Subfamily Rhysodinae Laporte, 1840

Tribe Clinidiini R.T. & J.R.Bell, 1978 *Clinidium* Kirby, 1830 *Grouvellina* R.T. & J.R.Bell, 1978 *Rhyzodiastes* Fairmaire, 1895 Tribe Dhysorini R.T. & J.R.Bell, 1978 *Dhysores* Grouvelle, 1903 *Neodhysores* R.T. & J.R.Bell, 1978 *Tangarona* R.T. & J.R.Bell, 1982 Tribe Leoglymmiini R.T. & J.R.Bell, 1978 *Leoglymmius* R.T. & J.R.Bell, 1978 Tribe Medisorini R.T. & J.R.Bell, 1987 *Medisores* R.T. & J.R.Bell, 1987 Tribe Omoglymmiini R.T. & J.R.Bell, 1978 *Arrowina* R.T. & J.R.Bell, 1978 *Omoglymmius* Ganglbauer, 1891 *Plesioglymmius* R.T. & J.R.Bell, 1978 *Shyrodes* Grouvelle, 1903 *Srimara* R.T. & J.R.Bell, 1978 *Xhosores* R.T. & J.R.Bell, 1978 *Yamatosa* R.T. & J.R.Bell, 1979 Tribe Rhysodini Laporte, 1840 *Kaveinga* R.T. & J.R.Bell, 1978 *Kupeus* R.T. & J.R.Bell, 1982 *Rhysodes* Germar, 1822 Tribe Sloanoglymmiini R.T. & J.R.Bell, 1991 *Sloanoglymmius* R.T. & J.R.Bell, 1991

### Subfamily Scaritinae Bonelli, 1810

Tribe Clivinini Rafinesque, 1815 Subtribe Androzelmina R.T.Bell, 1998 *Androzelma* Dostal, 1993 Subtribe Ardistomina Putzeys, 1867 *Ardistomis* Putzeys, 1846 *Aspidoglossa* Putzeys, 1846 *Kearophus* Dajoz, 2004 *Semiardistomis* Kult, 1950 Subtribe Clivinina Rafinesque, 1815 *Afrosyleter* Basilewsky, 1959 *Ancus* Putzeys, 1867 *Basilewskyana* Kult, 1959 *Brachypelus* Putzeys, 1867 *Cameroniola* Baehr, 2000 *Climax* Putzeys, 1861 *Clivina* Latreille, 1802 *Clivinarchus* Sloane, 1896 *Cryptomma* Putzeys, 1846 *Lachenus* Putzeys, 1846 *Leleuporella* Basilewsky, 1956 *Nannoryctes* Baehr, 2000 *Nyctosyles* Putzeys, 1867 *Orictites* Andrewes, 1931 *Platysphyrus* Sloane, 1905 *Psilidius* Jeannel, 1957 *Pyramoides* Bousquet, 2002 *Rhysocara* Sloane, 1916 *Rubidiclivina* Baehr, 2015 *Rugiluclivina* Balkenohl, 1996 *Semiclivina* Kult, 1947 *Sinesetosa* Balkenohl, 1996 *Syleter* Andrewes, 1941 *Thliboclivina* Kult, 1959 *Trilophidius* Jeannel, 1957 *Trilophus* Andrewes, 1927 *Trogloclivina* Deuve, 2003 *Whiteheadiana* Perrault, 1994 Subtribe Forcipatorina Bänninger, 1938 *Camptidius* Putzeys, 1867 *Camptodontus* Dejean, 1826 *Forcipator* Maindron, 1904 *Kultianella* Perrault, 1994 *Mesus* Chevrolat, 1858 *Obadius* Burmeister, 1875 *Oxygnathopsis* Louwerens, 1953 *Oxygnathus* Dejean, 1826 *Scolyptus* Putzeys, 1861 *Stratiotes* Putzeys, 1846 Subtribe Reicheiina Jeannel, 1957 *Alpiodytes* Jeannel, 1957 *Asioreicheia* Bulirsch & Magrini, 2014 *Catalanodytes* Sciaky, 1989 *Dalmatoreicheia* Magrini & Bulirsch, 2005 *Dimorphoreicheia* Magrini; Fancello & Leo, 2002 *Galicioreicheia* Felix & Bulirsch, 2015 *Guiodytes* Tian, 2013 *Gymnetoreicheia* Magrini; Fancello & Casale, 2019 *Iberodytes* Jeannel, 1949 *Ichnusodytes* Magrini; Fancello & Onnis, 2019 *Italodytes* G.Müller, 1938 *Kenyoreicheia* Bulirsch & Magrini, 2007 *Laoreicheia* Balkenohl, 2005 *Madagascareicheia* Magrini & Bulirsch, 2009 *Malagasyreicheia* Magrini; Bulirsch & Fancello, 2021 *Orientoreicheia* Bulirsch & Hurka, 1994 *Oxydrepanus* Putzeys, 1867 *Parareicheia* Jeannel, 1957 *Reicheadella* Reitter, 1913 *Reicheia* Saulcy, 1862 *Reicheidius* Jeannel, 1957 *Sikelioreicheia* Magrini & Fancello, 2019 *Spelaeodytes* L.Miller, 1863 *Typhloreicheia* Holdhaus, 1924 Subtribe Schizogeniina Dostal, 2017 *Baehrogenius* Dostal, 2017 *Coryza* Putzeys, 1867 *Halocoryza* Alluaud, 1919 *Lophocoryza* Alluaud, 1941 *Paracoryza* Basilewsky, 1952 *Psammocoryza* Hogan, 2006 *Schizogenius* Putzeys, 1846 Subtribe Sparostesina Dostal, 2017 *Bohemaniella* Bousquet, 2002 *Pseudoclivina* Kult, 1947 *Sparostes* Putzeys, 1867 Tribe Corintascarini Basilewsky, 1973 *Corintascaris* Basilewsky, 1952 Tribe Dyschiriini Kolbe, 1880 *Akephorus* LeConte, 1852 *Caledyschirius* Bulirsch, 2010 *Clivinopsis* Bedel, 1895 *Cribrodyschirius* Bruneau de Miré, 1952 *Dyschirius* Bonelli, 1810 *Neodyschirius* Kult, 1954 *Reicheiodes* Ganglbauer, 1891 *Setodyschirius* Fedorenko, 1996 *Striganoviella* Fedorenko, 2012 *Torretassoa* Schatzmayr & Koch, 1933 *Dyschiriomimus* Iablokoff-Khnzorian, 1960 Tribe Salcediini Alluaud, 1930 Subtribe Salcediina Alluaud, 1930 *Salcedia* Fairmaire, 1899 Subtribe Solenogenyina R.T.Bell, 1998 *Holoprizus* Putzeys, 1867 *Solenogenys* Westwood, 1859 Tribe Scaritini Bonelli, 1810 Subtribe Carenina W.J.MacLeay, 1887 *Carenidium* Chaudoir, 1868 *Carenum* Bonelli, 1813 *Epilectus* Blackburn, 1888 *Euryscaphus* W.J.MacLeay, 1865 *Laccopterum* W.J.MacLeay, 1878 *Monocentrum* Chaudoir, 1868 *Mouhotia* Laporte, 1862 *Neocarenum* Laporte, 1867 *Neoscaphus* Sloane, 1888 *Philoscaphus* W.J.MacLeay, 1871 *Scaraphites* Westwood, 1842 *Trichocarenum* Blackburn, 1892 Subtribe Pasimachina Putzeys, 1867 *Pasimachus* Bonelli, 1813 Subtribe Scapterina Putzeys, 1867 *Acanthoscelis* Dejean, 1825 *Oxylobus* Chaudoir, 1855 *Parathlibops* Basilewsky, 1958 *Passalidius* Chaudoir, 1863 *Scapterus* Dejean, 1826 *Steganomma* W.J.MacLeay, 1887 *Thlibops* Putzeys, 1867 Subtribe Scaritina Bonelli, 1810 *Anomophaenus* Fauvel, 1882 *Antilliscaris* Lorenz, 1998 *Baenningeria* Reichardt, 1976 *Coptolobus* Chaudoir, 1857 *Crepidopterus* Chaudoir, 1855 *Cryptoscaphus* Chaudoir, 1855 *Dinoscaris* Alluaud, 1902 *Distichus* Motschulsky, 1858 *Dyscaris* Bänninger, 1940 *Dyscherinus* Jeannel, 1955 *Dyscherus* Chaudoir, 1855 *Geoscaptus* Chaudoir, 1855 *Glyptogrus* Bates, 1875 *Gnaphon* Andrewes, 1920 *Haplogaster* Chaudoir, 1879 *Haplotrachelus* Chaudoir, 1855 *Macromorphus* Chaudoir, 1857 *Madascaris* Bänninger, 1938 *Mamboicus* Bates, 1886 *Mecynoscaris* Alluaud, 1930 *Menigius* Chaudoir, 1879 *Neochryopus* Bänninger, 1932 *Ochyropus* Schiödte, 1847 *Pachyodontus* Chaudoir, 1879 *Paradyscherus* Basilewsky, 1971 *Pilades* Heyne, 1895 *Prodyscherodes* Jeannel, 1955 *Prodyscherus* Jeannel, 1946 *Scarites* Fabricius, 1775 *Storthodontus* Chaudoir, 1855 *Tapinoscaris* Jeannel, 1946 *Tibioscarites* Bänninger, 1929 *Tonkinoscaris* Bänninger, 1956 *Typhloscaris* Kuntzen, 1914

### Subfamily Siagoninae Bonelli, 1813

Tribe Enceladini G.Horn, 1881 *Enceladus* Bonelli, 1813 Tribe Siagonini Bonelli, 1813 *Luperca* Laporte, 1840 *Siagona* Latreille, 1804

### Subfamily Trechinae Bonelli, 1810

Tribe Bembidarenini Maddison et al., 2019 *Andinodontis* Erwin & Toledano, 2010 *Argentinatachoides* Sallanave; Erwin & Roig-Juñent, 2008 *Bembidarenas* Erwin, 1972 *Tasmanitachoides* Erwin, 1972 Tribe Bembidiini Stephens, 1827 Subtribe Anillina Jeannel, 1937 *Afranillus* Giachino, 2015 *Afrodipnus* Giachino, 2015 *Angustanillus* Baehr & Main, 2016 *Anillaspis* Casey, 1918 *Anillinus* Casey, 1918 *Anillodes* Jeannel, 1963 *Anillopsidius* Coiffait, 1969 *Anillopsis* Jeannel, 1937 *Anillotarsus* Mateu, 1980 *Anillus* Jacquelin du Val, 1851 *Argiloborus* Jeannel, 1937 *Austranillus* Giachino, 2005 *Bafutyphlus* Bruneau de Miré, 1986 *Bhutanillus* M.E.Schmid, 1975 *Bulirschia* Giachino, 2008 *Bylibaraphanus* Giachino; Eberhard & Perina, 2021 *Caeconannus* Jeannel, 1963 *Caecoparvus* Jeannel, 1937 *Carayonites* Bruneau de Miré, 1986 *Cryptocharidius* M.Etonti & Mateu, 1992 *Cryptorites* Jeannel, 1950 *Dicropterus* Ehlers, 1883 *Elgonotyphlus* Sciaky & Zaballos, 1993 *Erwinanillus* Giachino; Eberhard & Perina, 2021 *Externanillus* Baehr & Main, 2016 *Geocharidius* Jeannel, 1963 *Geocharis* Ehlers, 1883 *Gilesdytes* Giachino; Eberhard & Perina, 2021 *Gracilanillus* Baehr & Main, 2016 *Gregorydytes* Giachino; Eberhard & Perina, 2021 *Hesperanillus* Baehr & Main, 2016 *Honduranillus* Zaballos, 1997 *Hygranillus* B.Moore, 1980 *Hypodipnites* Jeannel, 1963 *Hypotyphlus* Jeannel, 1937 *Iason* Giachino & Vailati, 2011 *Iberanillus* Español, 1971 *Illaphanus* W.J.MacLeay, 1865 *Kimberleytyphlus* Giachino; Eberhard & Perina, 2021 *Leleupanillus* Basilewsky, 1976 *Magnanillus* Baehr, 2017 *Malagasydipnus* Giachino, 2008 *Malagasytyphlus* Giachino, 2008 *Megastylulus* Giachino & Sciaky, 2003 *Mexanillus* Vigna Taglianti, 1973 *Microdipnites* Jeannel, 1957 *Microdipnodes* Basilewsky, 1960 *Microdipnus* Jeannel, 1937 *Microtyphlus* Linder, 1863 *Mystroceridius* Reichardt, 1972 *Neodipnus* Jeannel, 1957 *Neoillaphanus* Giachino; Eberhard & Perina, 2021 *Nesamblyops* Jeannel, 1937 *Nothanillus* Jeannel, 1962 *Orthotyphlus* Mateu & Zaballos, 1998 *Paranillopsis* Cicchino & Roig-Juñent, 2001 *Paranillus* Jeannel, 1949 *Parillaphanus* Baehr, 2018 *Parvocaecus* Coiffait, 1956 *Pelocharis* Jeannel, 1960 *Pelodiaetodes* B.Moore, 1980 *Pelodiaetus* Jeannel, 1937 *Pelonomites* Jeannel, 1963 *Perucharidius* Mateu & M.Etonti, 2002 *Pilbaradytes* Giachino; Eberhard & Perina, 2021 *Pilbaranillus* Baehr & Main, 2016 *Pilbaraphanus* Giachino; Eberhard & Perina, 2021 *Prioniomus* Jeannel, 1937 *Pseudanillus* Bedel, 1896 *Pseudillaphanus* Giachino, 2005 *Rhegmatobius* Jeannel, 1937 *Scotodipnus* Schaum, 1860 *Selenodipnus* Jeannel, 1963 *Serranillus* Barr, 1995 *Serratotyphlus* Giachino, 2008 *Stylulus* L.Schaufuss, 1882 *Tasmanillus* Giachino, 2005 *Typhlocharis* Dieck, 1869 *Typhlonesiotes* Jeannel, 1937 *Winklerites* Jeannel, 1937 *Zapotecanillus* Sokolov, 2013 *Zeanillus* Jeannel, 1937 *Zoianillus* Sciaky, 1994 Subtribe Bembidiina Stephens, 1827 *Amerizus* Chaudoir, 1868 *Asaphidion* Gozis, 1886 *Bembidion* Latreille, 1802 *Caecidium* Ueno, 1971 *Lionepha* Casey, 1918 *Ocys* Stephens, 1828 *Orzolina* Machado, 1987 *Sinechostictus* Motschulsky, 1864 Subtribe Horologionina Jeannel, 1949 *Horologion* Valentine, 1932 Subtribe Lovriciina Giachino; B.Gueorguiev & Vailati, 2011 *Lovricia* Pretner, 1979 *Neolovricia* Lakota; Jalzic & J.Moravec, 2009 *Paralovricia* Giachino; B.Gueorguiev & Vailati, 2011 Subtribe Tachyina Motschulsky, 1862 *Aenigmatachys* Baehr, 2020 *Afrotachys* Basilewsky, 1958 *Anomotachys* Jeannel, 1946 *Costitachys* Erwin, 1974 *Elaphropus* Motschulsky, 1839 *Kiwitachys* Larochelle & Larivière, 2007 *Liotachys* Bates, 1871 *Lymnastis* Motschulsky, 1862 *Meotachys* Erwin, 1974 *Micratopus* Casey, 1914 *Moirainpa* Erwin, 1984 *Nothoderis* Boyd & Erwin, 2016 *Pericompsus* LeConte, 1852 *Polyderidius* Jeannel, 1962 *Porotachys* Netolitzky, 1914 *Setitachys* Baehr, 2016 *Stigmatachys* Boyd & Erwin, 2016 *Straneoites* Basilewsky, 1947 *Tachys* Dejean, 1821 *Tachysbembix* Erwin, 2004 *Tachyta* Kirby, 1837 *Tachyxysta* Boyd & Erwin, 2016 Subtribe Xystosomina Erwin, 1994 *Erwiniana* Paulsen & A.Smith, 2003 *Geballusa* Erwin, 1994 *Gouleta* Erwin, 1994 *Inpa* Erwin, 1978 *Mioptachys* Bates, 1882 *Philipis* Erwin, 1994 *Xystosomus* Schaum, 1863 Tribe Pogonini Laporte, 1834 *Bedeliolus* Semenov, 1900 *Cardiaderus* Dejean, 1828 *Diodercarus* Lutshnik, 1931 *Diplochaetus* Chaudoir, 1872 *Ochtozetus* Chaudoir, 1872 *Olegius* Komarov, 1996 *Pogonistes* Chaudoir, 1872 *Pogonopsis* Bedel, 1898 *Pogonus* Dejean, 1821 *Sirdenus* Dejean, 1828 *Syrdenoidius* Baehr & Hudson, 2001 *Thalassotrechus* Van Dyke, 1918 Tribe Sinozolini Deuve, 1997 *Chaltenia* Roig-Juñent & Cicchino, 2001 *Phrypeus* Casey, 1924 *Sinozolus* Deuve, 1997 Tribe Trechini Bonelli, 1810 Subtribe Aepina Fowler, 1887 *Aepopsis* Jeannel, 1922 *Aepus* Leach in Samouelle, 1819 *Kenodactylus* Broun, 1909 *Kiwitrechus* Larochelle & Larivière, 2007 *Maoritrechus* Brookes, 1932 *Neanops* Britton, 1962 *Oarotrechus* Townsend, 2010 *Temnostega* Enderlein, 1905 *Thalassobius* Solier, 1849 *Waiputrechus* Townsend, 2010 Subtribe Perileptina Sloane, 1903 *Apoplotrechus* Alluaud, 1915 *Neoblemus* Jeannel, 1923 *Perileptus* Schaum, 1860 Subtribe Trechina Bonelli, 1810 *Accoella* Ueno, 1990 *Acheroniotes* Lohaj & Lakota, 2010 *Adriaphaenops* Noesske, 1928 *Aepiblemus* Belousov & Kabak, 1993 *Agonotrechus* Jeannel, 1923 *Agostinia* Jeannel, 1928 *Alanorites* Belousov, 1998 *Albanotrechus* Casale & V.B.Gueorguiev, 1994 *Allegrettia* Jeannel, 1928 *Allotrechiama* Ueno, 1970 *Ameroduvalius* Valentine, 1952 *Anchotrechus* Jeannel, 1927 *Andinorites* Mateu & Belles, 1980 *Andinotrechus* Mateu, 1981 *Anillidius* Jeannel, 1928 *Anophthalmus* Sturm, 1844 *Aphaenopidius* G.Müller, 1913 *Aphaenops* Bonvouloir, 1862 *Aphaenopsis* G.Müller, 1913 *Apocimmerites* Belousov, 1998 *Apoduvalius* Jeannel, 1953 *Aputrechisibus* Trezzi, 2007 *Arctaphaenops* Meixner, 1925 *Aspidaphaenops* Ueno, 2006 *Austrotrechus* B.Moore, 1972 *Awatrechus* Ueno, 1955 *Balazucellus* Deuve, 2001 *Bathytrechus* Ueno, 2005 *Beronaphaenops* B.V.Gueorguiev, 2012 *Birmaphaenops* Deuve, 2017 *Blemus* Dejean, 1821 *Boldoriella* Jeannel, 1928 *Boreaphaenops* Ueno, 2002 *Bothynotrechus* B.Moore, 1972 *Casaleaphaenops* Tian; Huang & Ma, 2021 *Cathaiaphaenops* Deuve, 1996 *Caucasaphaenops* Belousov, 1999 *Caucasorites* Belousov & Zamotajlov, 1997 *Chaetoduvalius* Jeannel, 1928 *Chaetotrechiama* Ueno, 1982 *Chiapadytes* Vigna Taglianti, 1977 *Chu* Tian & He, 2020 *Cimbrodytes* Piva, 2021 *Cimmerites* Jeannel, 1928 *Cimmeritodes* Deuve, 1996 *Ckacus* Tian & Huang, 2021 *Columbitrechus* Mateu, 1982 *Coreoblemus* Ueno, 1969 *Croatotrechus* Casale & Jalzic, 1999 *Dabatrechus* Ueno, 2004 *Dactylotrechus* Belousov & Kabak, 2003 *Daiconotrechus* Ueno, 1971 *Dalmataphaenops* Monguzzi, 1993 *Darlingtonea* Valentine, 1952 *Derossiella* Queinnec, 2008 *Deuveaphaenops* Tian & Huang, 2017 *Dianotrechus* Tian, 2016 *Doderotrechus* Vigna Taglianti, 1968 *Dongoblemus* Deuve & Tian, 2016 *Dongodytes* Deuve, 1993 *Dracotrechus* Ueno, 2010 *Duvalioblemus* Deuve, 1995 *Duvaliomimus* Jeannel, 1928 *Duvaliopsis* Jeannel, 1928 *Duvalius* Delarouzée, 1859 *Eocnides* Jeannel, 1954 *Epaphiopsis* Ueno, 1953 *Epaphiotrechus* Deuve & Kavanaugh, 2016 *Epaphius* Leach in Samouelle, 1819 *Erebotrechus* Britton, 1964 *Escolatrechus* Mateu, 2003 *Eutrechopsis* B.Moore, 1972 *Eutrechus* B.Moore, 1972 *Geotrechus* Jeannel, 1919 *Giraffaphaenops* Deuve, 2002 *Glabroduvalius* Vrbica; Curcic; Antic & Curcic, 2013 *Goedetrechus* B.Moore, 1972 *Gotoblemus* Ueno, 1970 *Graciliblemus* Deuve & Tian, 2016 *Guatemalatrechus* Trezzi, 2003 *Guiaphaenops* Deuve, 2002 *Guizhaphaenops* Vigna Taglianti, 1997 *Guizhaphaenopsoides* Tian; Huang & Li, 2021 *Gulaphaenops* Ueno, 1987 *Himalaphaenops* Ueno, 1980 *Himalotrechus* Belousov; Kabak & J.Schmidt, 2019 *Homaloderodes* Jeannel, 1962 *Huoyanodytes* Tian & Huang, 2016 *Iberotrechus* Jeannel, 1920 *Iga* Ueno, 1953 *Incatrechus* Mateu & Belles, 1982 *Inotrechus* Dolzhanski & Ljovushkin, 1989 *Ishidatrechus* Ueno, 1956 *Ishikawatrechus* Habu, 1950 *Italaphaenops* Ghidini, 1964 *Iyotrechus* Ueno & Naito, 2009 *Jalzicaphaenops* Lohaj & Lakota, 2010 *Jeannelius* Kurnakov, 1959 *Jiangxiaphaenops* Ueno & Clarke, 2007 *Jiulongotrechus* Tian; Huang & Wang, 2015 *Junaphaenops* Ueno, 1997 *Junnanotrechus* Ueno & Yin, 1993 *Kettlotrechus* Townsend, 2010 *Kosswigia* Jeannel, 1947 *Kozlovites* Jeannel, 1935 *Kupetrechus* Larochelle & Larivière, 2007 *Kurasawatrechus* Yoshida & Nomura, 1952 *Kusumia* Ueno, 1952 *Lanxangaphaenops* Deuve, 2012 *Laoblemus* Ueno, 2006 *Laosaphaenops* Deuve, 2000 *Lessinodytes* Vigna Taglianti, 1982 *Libotrechus* Ueno, 1998 *Luoxiaotrechus* Tian & Yin, 2013 *Luyatrechus* M.Etonti & Mateu, 2000 *Luzonotrechus* Ueno, 1979 *Mamesdytes* Trezzi, 2003 *Masuzoa* Ueno, 1960 *Masuzonoblemus* Ueno, 1989 *Mayaphaenops* Vigna Taglianti, 1977 *Meganophthalmus* Kurnakov, 1959 *Mexaphaenops* Bolivar y Pieltain, 1942 *Mexitrechus* Barr, 1982 *Miaotrechus* Tian; Chen & Ma, 2020 *Microblemus* Ueno, 2007 *Mimanillus* B.Moore, 1972 *Mimotrechus* B.Moore, 1972 *Minimaphaenops* Deuve, 2000 *Minosaphaenops* Queinnec, 2008 *Minutotrechus* Deuve & Kavanaugh, 2016 *Nannotrechus* Winkler, 1926 *Nanotrechiama* Belousov & Kabak, 2018 *Neaphaenops* Jeannel, 1920 *Nelsonites* Valentine, 1952 *Neotrechus* G.Müller, 1913 *Nesaeoduvalius* Casale & Giachino, 2016 *Nesiotrechus* Ueno, 1995 *Nipponaphaenops* Ueno, 1971 *Nototrechus* B.Moore, 1972 *Nunbergites* Pawlowski & Stachowiak, 1991 *Omalodera* Blanchard, 1842 *Oodinotrechus* Ueno, 1998 *Oroblemites* Ueno & Pawlowski, 1981 *Oroblemus* Ueno & Yoshida, 1966 *Orotrechus* G.Müller, 1913 *Oxytrechus* Jeannel, 1927 *Panaphaenops* Tian; Huang & Ma, 2021 *Paracimmerites* Belousov, 1998 *Paraphaenops* Jeannel, 1916 *Paratrechiotes* Ueno, 1995 *Paratrechus* Jeannel, 1920 *Petraphaenops* Delic; Kapla & Colla, 2019 *Pheggomisetes* Knirsch, 1923 *Pilosaphaenops* Deuve & Tian, 2008 *Plesioaphaenops* Deuve & Tian, 2011 *Pogonoschema* Jeannel, 1927 *Pontodytes* Casale & Giachino, 1989 *Porocimmerites* Belousov, 1998 *Protrechiama* Belousov & Kabak, 2003 *Pseudanophthalmus* Jeannel, 1920 *Pseudaphaenops* Winkler, 1912 *Pseudocnides* Jeannel, 1927 *Pseudotrechisibus* Mateu & Belles, 1982 *Puertrechus* Belousov & Kabak, 2014 *Putzeysius* Jeannel, 1962 *Qianaphaenops* Ueno, 2000 *Qianotrechus* Ueno, 2000 *Queinnectrechus* Deuve, 1992 *Rakantrechus* Ueno, 1951 *Ryugadous* Habu, 1950 *Sardaphaenops* Cerruti & Henrot, 1956 *Satotrechus* Ueno, 2006 *Sbordoniella* Vigna Taglianti, 1980 *Scotoplanetes* Absolon, 1913 *Scototrechus* Britton, 1962 *Shenaphaenops* Ueno, 1999 *Shennongotrechus* Ueno, 2004 *Shenoblemus* Tian & Fang, 2020 *Shilinotrechus* Ueno, 2003 *Shiqianaphaenops* Tian, 2016 *Shuangheaphaenops* Tian, 2017 *Shuaphaenops* Ueno, 1999 *Sichuanotrechus* Deuve, 2005 *Sidublemus* Tian & Yin, 2013 *Sinaphaenops* Ueno & Wang, 1991 *Sinocimmerites* Deuve, 2007 *Sinotrechiama* Ueno, 2000 *Sinotroglodytes* Deuve, 1996 *Sloanella* Jeannel, 1927 *Speotrechus* Jeannel, 1922 *Stevensius* Jeannel, 1923 *Stygiotrechus* Ueno, 1958 *Subilsia* Español, 1967 *Sumatrechus* Deuve, 2005 *Superbotrechus* Deuve & Tian, 2009 *Suzuka* Ueno, 1956 *Taiwanotrechus* Ueno, 1987 *Taniatrechus* Belousov & Dolzhanski, 1994 *Tasmanorites* Jeannel, 1927 *Tasmanotrechus* B.Moore, 1972 *Taurocimmerites* Belousov, 1998 *Thalassoduvalius* Ueno, 1956 *Tianeotrechus* Tian & Tang, 2016 *Tianzhuaphaenops* Zhao & Tian, 2016 *Tienmutrechus* Suenson, 1957 *Tonkinaphaenops* Deuve, 2013 *Toshiaphaenops* Ueno, 1999 *Trechepaphiama* Deuve & Kavanaugh, 2016 *Trechepaphiopsis* Deuve & Kavanaugh, 2016 *Trechiama* Jeannel, 1927 *Trechiamiotes* Deuve, 1998 *Trechicomimus* Mateu & Nègre, 1972 *Trechiella* Jeannel, 1927 *Trechimorphus* Jeannel, 1927 *Trechinotus* Jeannel, 1962 *Trechiotes* Jeannel, 1954 *Trechisibus* Motschulsky, 1862 *Trechistus* B.Moore, 1972 *Trechoblemus* Ganglbauer, 1891 *Trechus* Clairville, 1806 *Trichaphaenops* Jeannel, 1916 *Troglocimmerites* Ljovuschkin, 1970 *Tropidotrechus* Jeannel, 1927 *Typhlotrechus* G.Müller, 1913 *Uenoaphaenops* Tian & He, 2020 *Uenoites* Belousov & Kabak, 2016 *Uenotrechus* Deuve & Tian, 1999 *Ushijimaella* Ueno, 1980 *Velebitaphaenops* Casale & Jalzic, 2012 *Velesaphaenops* S.Curcic et al., 2018 *Vietotrechus* Ueno, 1995 *Wanhuaphaenops* Tian & Wang, 2016 *Wanoblemus* Tian & Fang, 2016 *Wulongoblemus* Ueno, 2007 *Xenotrechus* Barr & Krekeler, 1967 *Xiangxius* Tian; Huang & Li, 2021 *Xuedytes* Tian & Huang, 2017 *Yalongaphaenops* Belousov & Kabak, 2021 *Yamautidius* Ueno, 1957 *Yanzaphaenops* Ueno, 2010 *Yunotrechus* Tian & Huang, 2014 *Zhijinaphaenops* Ueno & Ran, 2002 Subtribe Trechodina Jeannel, 1926 *Amblystogenium* Enderlein, 1905 *Canarobius* Machado, 1987 *Cnides* Motschulsky, 1862 *Cothresia* Jeannel, 1964 *Cyphotrechodes* Jeannel, 1926 *Eotrechodes* Ueno; Lafer & Sundukov, 1995 *Himalotrechodes* Ueno, 1981 *Iberotrechodes* Faille et al., 2021 *Pachydesus* Motschulsky, 1864 *Pachytrechodes* Jeannel, 1960 *Paratrechodes* Jeannel, 1926 *Scaurotrechodes* Geginat, 2006 *Spelaeovulcania* Machado, 1987 *Sporades* Fauvel, 1882 *Thalassophilus* Wollaston, 1854 *Trechobembix* Jeannel, 1926 *Trechodes* Blackburn, 1901 *Trechosia* Jeannel, 1926 *Trechosiella* Jeannel, 1960 *Antarctotrechus* Ashworth & Erwin, 2016 *Trechinites* Heer, 1862 *Trechoides* Motschulsky, 1856 Tribe Zolini Sharp, 1886 Subtribe Merizodontina Sloane, 1920 *Idacarabus* Lea, 1910 *Maungazolus* Larochelle & Larivière, 2017 *Merizodus* Solier, 1849 *Percodermus* Sloane, 1920 *Pseudoopterus* Csiki, 1928 *Pterocyrtus* Sloane, 1920 *Sloaneana* Csiki, 1933 *Synteratus* Broun, 1909 *Thayerella* Baehr, 2016 Subtribe Zolina Sharp, 1886 *Oopterus* Guérin-Méneville, 1841 *Zolus* Sharp, 1886

### Subfamily Xenaroswellianinae Erwin, 2007

*Xenaroswelliana* Erwin, 2007

### Subfamily †Conjunctiinae Ponomarenko, 1977

*Conjunctia* Ponomarenko, 1977

### Subfamily †Protorabinae Ponomarenko, 1977

*Aethocarabus* Ren; Lu & Guo, 1995 *Atrirabus* Hong & Wang, 1990 *Cordorabus* Ponomarenko, 1977 *Cretorabus* Ponomarenko, 1977 *Denudirabus* Ren; Lu & Guo, 1995 *Lirabus* Hong, 1992 *Lithorabus* Ponomarenko, 1977 *Magnirabus* Hong & Wang, 1990 *Mesorabus* Ponomarenko, 1977 *Nebrorabus* Ponomarenko, 1989 *Ovrabites* Ponomarenko, 1977 *Penecupes* Ren, 1995 *Protorabus* Ponomarenko, 1977 Not Assigned to Subfamily *Agatoides* Motschulsky, 1856 *Amphoxyne* Bode, 1953 *Carabites* Heer, 1852 *Cavicarabus* Hong, 1991 *Conexicoxa* Lin, 1986 *Cymatopterus* Lomnicki, 1894 *Fangshania* Hong, 1981 *Glenopterus* Heer, 1847 *Hebeicarabus* Hong, 1983 *Megacarabus* Hong, 1983 *Meileyingia* Hong, 1987 *Miocarabus* Hong, 1983 *Neothanes* Scudder, 1890 *Procarabus* Oppenheim, 1888 *Prosynactus* Bode, 1953 *Shanwangicarabus* Hong, 1985 *Sinis* Heer, 1862 *Sinocalosoma* Hong & Wang, 1986 *Sinocaralosoma* Hong, 1984 *Sunocarabus* Hong, 1987 *Tauredon* Handlirsch, 1910 *Wuchangicarabus* Hong, 1991 *Xishanocarabus* Hong, 1984 *Yunnanocarabus* Lin, 1977


title: Tony Steedman

abstract: Anthony Francis Steedman was an English character actor, perhaps best known for roles in British TV drama series of the 1970s and 1980s, and for his role as Socrates in Bill & Ted's Excellent Adventure.

text:
# Tony Steedman

**Anthony Francis Steedman** (21 August 1927  4 February 2001) was an English character actor, perhaps best known for roles in British TV drama series of the 1970s and 1980s, and for his role as Socrates in *Bill & Ted's Excellent Adventure*.

## Career

In addition to film work, his theatre appearances were extensive, including Ray Cooney's comedy *Her Royal Highness* in the West End of London, and he appeared in numerous television productions, including episodes of *The Feathered Serpent*, *Thriller*, *The Professionals*, *Randall and Hopkirk (Deceased)*, *Coronation Street*, *The Rivals of Sherlock Holmes*, *The Avengers*, *Babylon 5*, *The Sweeney*, *Minder* and *Barry Morse presents Strange But True*. He had a recurring role in the British soap opera Crossroads as Dr. Butterworth. He was also a regular in the final two series of BBC sitcom *Citizen Smith* (1979–80). He holds the distinction of portraying Nazi Gen. Alfred Jodl twice, first in *The Death of Adolf Hitler* (1973) and later in *The Bunker* (1981). Following his role in *Bill and Ted*, he voiced the character of Justin Hammer in the first season of *Iron Man*.

### Partial filmography

## Personal life

Steedman met his first wife, Ann Taylor, in 1957 during a production of *Oklahoma!* and they were married in November of that year in Brackley. They had one daughter, and divorced in 1963.

Steedman's second wife was actress Judy Parfitt, whom he met in 1960 at the Birmingham Rep. They were married in 1963 and jointly ran Allerway Limited, an artistic creation company. They had one son together.

Steedman died on 4 February 2001.

## InfoBox

| Tony Steedman | |
| --- | --- |
| Publicity still, 1984 | |
| Born | Anthony Francis Steedman
(1927-08-21)21 August 1927
Warwickshire, England |
| Died | 4 February 2001(2001-02-04) (aged 73) |
| Resting place | East London Cemetery, Plaistow, London |
| Occupation | Actor |
| Years active | 1957–1998 |
| Spouses | *   Ann Taylor  (m. 1957; div. 1963) *   Judy Parfitt (m. 1963) |
| Children | 2 |


title: Stone Age Complication

abstract: Stone Age Complication is an EP by Queens of the Stone Age containing six B-sides and covers.

text:
# Stone Age Complication

Professional ratings
| Review scores | |
| --- | --- |
| Source | Rating |
| AllMusic |  |

***Stone Age Complication*** is an EP by Queens of the Stone Age containing six B-sides and covers.

## Track listing

Lead vocals by Josh Homme, except where noted.
| No. | Title | Writer(s) | Lead vocals | Length |
| --- | --- | --- | --- | --- |
| 1. | "Who'll Be the Next in Line" (The Kinks cover) | Ray Davies | Nick Oliveri | 2:31 |
| 2. | "Wake Up Screaming" (Subhumans cover) | Subhumans | Oliveri | 5:03 |
| 3. | "No One Knows (UNKLE Remix)" | Josh Homme, Mark Lanegan |  | 4:37 |
| 4. | "Most Exalted Potentate of Love" (The Cramps cover) | The Cramps |  | 2:46 |
| 5. | "Born to Hula" (2000 version) | Homme |  | 5:55 |
| 6. | "The Bronze" | Homme |  | 3:41 |
| Total length: | | | | 24:33 |

## InfoBox

| Stone Age Complication | |
| --- | --- |
|  | |
| EP by Queens of the Stone Age | |
| Released | October 4, 2004 (2004-10-04) |
| Recorded | 1998–2002 |
| Genre | Hard rock |
| Length | 24:33 |
| Label | Interscope |
| Producer | Queens of the Stone Age |
| Queens of the Stone Age chronology | |
| *Songs for the Deaf*
(2002) ***Stone Age Complication***
(2004) *Lullabies to Paralyze*
(2005) | |


title: Westhead

abstract: Westhead is a village in the West Lancashire district of Lancashire, England. As of 2014, the estimated population was 886.

text:
# Westhead

**Westhead** is a village in the West Lancashire district of Lancashire, England. As of 2014, the estimated population was 886.

## History

Westhead was, for most of its existence, a hamlet in the township of Lathom. It was located in the West Derby Hundred. In 1066, the Lord of the area was Uhtred.

In 1851, Westhead Church was built with land donated from Edward Wilbraham-Bootle, 1st Baron Skelmersdale, the cost of which was paid for by the Earl of Derby and the sandstone was donated by Edward Stanley of Cross Hall, who owned the Ruff Wood quarry in Lathom at the time.

Westhead Halt was a request stop on the Skelmersdale to Rainford line, and evidence of the former line can be seen around the town.

### Toponymy

Westhead means "western hill" and comes from the Old English *west* \+ *hēafod*. The name was recorded as *Westheft* c. 1190 and *Westheved* c. 1200, and refers to the western headland at the end of the ridge that leads towards Scarth Hill.

## Geography

Westhead is located in rural West Lancashire and is part of the unparished area of Ormskirk. It lies about 1.2 miles (1.9 km) east of Ormksirk town centre and is bordered to the north, east and south by the civil parishes of Burscough, Lathom, Lathom South and Bickerstaffe.

Westhead is about 15 miles (24 km) southwest of Preston, the administrative centre of Lancashire, and 182 miles (293 km) northwest of London.

## Culture and community

Westhead Carnival is a popular summertime fête which has been held annually since 1973. The event is typically held over three days at the start of July and hosts a variety of activities, including a carnival parade, funfair rides, car boot sale, bingo and fancy dress ball. In 2014, the carnival was attended by around 3,000 people and raised over £2,000 for local community projects.

## Transport

The A577 road runs east through Westhead for a distance of approximately 1 mile (1.6 km), connecting the nearby towns of Ormskirk and Skelmersdale. There are several bus services operated by Arriva North West that pass along this route: the 310 is a half-hourly service between Skelmersdale and Liverpool, while the 375 and 385 are hourly services (alternating every half hour) from Southport to Wigan. The nearest motorway link is junction 3 of the M58, about 1.7 miles (2.7 km) to the south at Bickerstaffe.

Westhead Halt, situated on Castle Lane, was a former railway station on the Skelmersdale Branch which connected Ormskirk with Rainford. However, the station closed in 1951 and has since been demolished, the line having closed to all traffic in 1963. The nearest railway station today is Ormskirk, about 1.2 miles (1.9 km) east, which provides passenger services to Liverpool and Preston.

## Education

Westhead Lathom St James is a Church of England primary school associated with the nearby parish church of St James. The school is located on School Lane near the village centre and provides mixed gender education for children aged four to eleven. An Ofsted report published in January 2015 gave Westhead Lathom St James an "Outstanding" rating in all categories, and in December 2015 it was ranked amongst the top 1000 primary schools in England in the National Curriculum assessment performance tables. As of September 2019, the headteacher was Miss Helen Clark.

The school building with attached master's house dates from 1889 and is a Grade II listed building. It is made of sandstone with crazy paving walling and a slate roof, and is built in the Gothic style. The school has a single storey with a T-shaped plan, formed by a main range and cross wing to the left, while the house has two storeys and is attached to the right.

## Religious sites

St James Church is a Grade II listed building situated on Vicarage Lane, about 0.5 miles (0.8 km) southwest of the village. The ecclesiastical parish of St James Westhead is in the Anglican Diocese of Liverpool, the archdeaconry of Ormskirk, and forms part of a benefice with the nearby parish of Christ Church, Newburgh. In the churchyard are two Commonwealth War Graves, one belonging to a soldier of World War I and the other an airman of World War II.

## InfoBox

| Westhead | |
| --- | --- |
| Village | |
| Wigan Road, Westhead | |
| WestheadShown within West LancashireWestheadLocation within Lancashire | |
| Area | 0.203 km2 (0.078 sq mi) |
| Population | 886 (2014 estimate) |
| Density | 4,365/km2 (11,310/sq mi) |
| OS grid reference | SD441077 |
| London | 182 mi (293 km) SE |
| District |  |
| Shire county |  |
| Region |  |
| Country | England |
| Sovereign state | United Kingdom |
| Post town | Ormskirk |
| Postcode district | L40 |
| Dialling code | 01695 |
|  | |
| Police | Lancashire |
| Fire | Lancashire |
| Ambulance | North West |
|  | |
| UK Parliament |  |
|


title: William J. Livsey

abstract: William James "Lipp" Livsey was a United States Army general who served as the Commander in Chief of United Nations Command/Commander in Chief, ROK/U.S. Combined Forces Command/Commander, United States Forces Korea/Commanding General, Eighth United States Army from 1984 to 1987.

text:
# William J. Livsey

**William James "Lipp" Livsey** (June 8, 1931 – June 18, 2016) was a United States Army general who served as the Commander in Chief of United Nations Command/Commander in Chief, ROK/U.S. Combined Forces Command/Commander, United States Forces Korea/Commanding General, Eighth United States Army from 1984 to 1987.

## Early life and education

Livsey was born on June 8, 1931, in Clarkston, Georgia. He is a 1952 graduate of North Georgia College. Years later, he served as a trustee for the university foundation. He received his commission as a second lieutenant of Infantry in the United States Army. He has a master's degree in Psychology from Vanderbilt University. His military schooling includes the Infantry Officer Basic and Advanced Courses, the United States Army Command and General Staff College, where he graduated first in his class, the Armed Forces Staff College, and the Army War College.

## Career

In 1953, Livsey served his first combat tour as platoon leader with the 3rd Infantry Division in South Korea. He commanded his first company with the 30th Infantry at Fort Benning, Georgia in 1954. From 1958 to 1961, he served in Germany as a company commander in the 2nd Armored Rifle Battalion, 36th Infantry, 3rd Armored Division. He served his second combat tour in Vietnam from 1967 to 1968 with the 4th Infantry Division as Assistant Chief of Staff, G-3 (Operations), and as the Commander, 2nd Battalion, 35th Infantry of the 3rd Brigade Task Force.

In May 1971, Livsey took command of the 2nd Brigade 4th Infantry Division (Mechanized) at Fort Carson. He later returned to Fort Carson in 1974 to serve as the Assistant Division Commander (Support) and as the Assistant Division Commander (Maneuver) for the 4th Infantry Division until June 1976. From July 1977 to May 1979, he was the Commanding General of the Infantry Center and Commandant of the United States Army Infantry School at Fort Benning, Georgia.

In May 1979, Livsey became the Commanding General of the 8th Infantry Division (Mechanized), United States Army Europe where he served until June 1981. He then assumed the position of Commanding General, VII Corps, USAREUR, and served there until July 1983. From July 1983 until April 1984 he served as the Deputy Commanding General, United States Army Forces Command as well as Commanding General of the Third United States Army, Fort McPherson, Georgia.

Other notable assignments include, I Corps Chief of Staff in Korea, Executive to the Army Chief of Staff in Washington, D.C., and Southeast Asia, staff officer in the Office of Research and Development, and leadership instructor at West Point.

In May 1984, and until his retirement on June 30, 1987, Livsey served as Commander-in-Chief, United Nations Command/Commander-in-Chief Combined Forces Command/Commander, United States Forces, Korea/Commanding General Eighth United States Army.

## Awards and decorations

Livsey's awards and decorations include the Defense Distinguished Service Medal, Army Distinguished Service Medal (with oak leaf cluster), Silver Star, Legion of Merit (with 3 oak leaf cluster), Distinguished Flying Cross, Bronze Star Medal with "V" device, Air Medals, Army Commendation Medal with "V" device (with 2 oak leaf cluster), Combat Infantryman Badge (second award), Army Staff Identification Badge, and the Parachutist Badge.

## Personal life

In October 2008, a portion of Highway 314 in Fayetteville, Georgia was renamed as the "General Bill Livsey Highway" in Livsey's honor. Numerous active and retired military leaders, including Major General Jack Wheeler, were in attendance. Also in attendance was HeroBox, a non-profit organization that sends custom care packages to American soldiers.

On April 18, 2009, the drill field at the center of the University of North Georgia's campus was renamed the William J. "Lipp" Livsey Drill Field in Livsey's honor. A commemorative plaque bearing Livsey's likeness was added near the reviewing stand.

On August 18, 2015, Livsey was arrested for allegedly assaulting Ryan Irvin, a food delivery man for a local Chinese restaurant, and resisting arrest in Fayetteville, Georgia. Meanwhile, the restaurant wanted to drop the incident entirely. He died on June 18, 2016, at the age of 85.

This article incorporates public domain material from the United States Army

## InfoBox

| William J. Livsey | |
| --- | --- |
| General William J. Livsey | |
| Nickname(s) | "Lipp" |
| Born | (1931-06-08)June 8, 1931
Clarkston, Georgia |
| Died | June 18, 2016(2016-06-18) (aged 85)
Fayetteville, Georgia |
| Allegiance | United States |
| Service/branch | United States Army |
| Years of service | 1952–1987 |
| Rank | General |
| Commands | Eighth United States Army
Third United States Army
8th Infantry Division
United States Army Infantry School
2nd Brigade, 4th Infantry Division
2nd Battalion, 35th Infantry Regiment |
| Battles/wars | Korean War
Vietnam War |
| Awards | Defense Distinguished Service Medal
Army Distinguished Service Medal (2)
Silver Star
Legion of Merit (4)
Distinguished Flying Cross
Bronze Star Medal |
| Spouse(s) | Bena Sue Burns |


title: Negratín Reservoir

abstract: Negratín Reservoir is a reservoir located in Freila, province of Granada, Andalusia, Spain and it is considered one of 11 other reservoirs in Granada.

text:
# Negratín Reservoir

**Negratín Reservoir** is a reservoir located in Freila, province of Granada, Andalusia, Spain and it is considered one of 11 other reservoirs in Granada.

## Introduction

Lake Negratin stands as one of the largest freshwater lakes in Andalusia, created by the Negratin Dam along the Guadiana Menor River. Accessible via the A-315, it can be reached from the nearby villages of Freila, Zujar, or Cuevas del Campo. The lake presents a stunning vista, displaying hues ranging from aquamarine to cobalt blue, influenced by the time of day and the sun's position. Framed by rugged cliffs on its northern bank, the lake provides a picturesque retreat for those seeking tranquility amid natural beauty.

For water enthusiasts, activities such as canoeing and sailing are available through the El Negratin Club Nautico. This sailing club, complete with a lakeside restaurant, is conveniently situated and marked from the village of Cuevas del Campo, offering a delightful opportunity to engage in water sports while enjoying the serene surroundings.

## Dam structure

Mixed gravity and loose materials with asphalt screen, straight plant with a crest length of 439 m and 10 meters width. Gated spillway. Botton drains type with 2 channels and a drainage capacity of 120 m3/s.

## Characteristics

Covering an expanse of 2,170 hectares and boasting a volumetric capacity of 567 cubic hectometers, this reservoir ranks as the fourth largest in the autonomous community of Andalusia.

The dam is a composite structure, combining elements of both gravity and loose material construction, featuring an asphalt lining for waterproofing. This particular design was opted for due to the insufficient stability of the left abutment, preventing the construction of a concrete dam in that location.

**Main Objectives-Purpose of the Negratin Dam & Reservoir:**

* Irrigation * Electricity * Fishing * Ecological Flow

## InfoBox

| Negratín Reservoir | |
| --- | --- |
| Embalse del Negratín from Baza | |
| Negratín Reservoir | |
| Location | Freila |
| Coordinates | 37°34′20″N 2°53′16″W / 37.57222°N 2.88778°W / 37.57222; -2.88778 |
| Type | reservoir |
| Primary inflows | Guadiana Menor River |
| Basin countries | Spain |
| Built | 1984 (1984) |


title: Semicircle law (quantum Hall effect)

abstract: The semicircle law, in condensed matter physics, is a mathematical relationship that occurs between quantities measured in the quantum Hall effect. It describes a relationship between the anisotropic and isotropic components of the macroscopic conductivity tensor σ, and, when plotted, appears as a semicircle. The semicircle law was first described theoretically in Dykhne and Ruzin's analysis of the quantum Hall effect as a mixture of 2 phases: a free electron gas, and a free hole gas. Mathematically, it states that σ x x 2 + 2 = 2 where σ is the mean-field Hall conductivity, and σ⁰ is a parameter that encodes the classical conductivity of each phase. A similar law also holds for the resistivity. A convenient reformulation of the law mixes conductivity and resistivity: σ x y 0 = ρ x y 0 2 + 2 = e 2 h where n is an integer, the Hall divisor. Although Dykhne and Ruzin's original analysis assumed little scattering, an assumption that proved empirically unsound, the law holds in the coherent-transport limits commonly observed in experiment. Theoretically, the semicircle law originates from a representation of the modular group Γ0(2), which describes a symmetry between different Hall phases. That group's strong connections to number theory also appear: Hall phase transitions exhibit a selection rule | p q ′ − p ′ q | = 1 that also governs the Farey sequence. Indeed, plots of the semicircle law are also Farey diagrams. In striped quantum Hall phases, the relationship is slightly more complex, because of the broken symmetry: { σ 1 σ 2 + 2 = 2 σ h 0 = e 2 / h Here σ1 and σ2 describe the macroscopic conductivity in directions aligned with and perpendicular to the stripes.

text:
# Semicircle law (quantum Hall effect)

The **semicircle law**, in condensed matter physics, is a mathematical relationship that occurs between quantities measured in the quantum Hall effect. It describes a relationship between the anisotropic and isotropic components of the macroscopic conductivity tensor σ, and, when plotted, appears as a semicircle.

The semicircle law was first described theoretically in Dykhne and Ruzin's analysis of the quantum Hall effect as a mixture of 2 phases: a free electron gas, and a free hole gas. Mathematically, it states that $\sigma \_{xx}^{2}+(\sigma \_{xy}-\sigma \_{xy}^{0})^{2}=(\sigma \_{xx}^{0})^{2}$where σ is the mean-field Hall conductivity, and *σ*0 is a parameter that encodes the classical conductivity of each phase. A similar law also holds for the resistivity.

A convenient reformulation of the law mixes conductivity and resistivity: $\sigma \_{xy}^{0}={\frac {\rho \_{xy}^{0}{(\rho \_{xy}^{0})^{2}+(\rho \_{xx}^{0})^{2}={\frac {e^{2}{h}\left(n+{\frac {1}{2}\right)}}}}}}$where n is an integer, the Hall divisor.

Although Dykhne and Ruzin's original analysis assumed little scattering, an assumption that proved empirically unsound, the law holds in the coherent-transport limits commonly observed in experiment.

Theoretically, the semicircle law originates from a representation of the modular group Γ0(2), which describes a symmetry between different Hall phases. (Note that this is not a symmetry in the conventional sense; there is no conserved current.) That group's strong connections to number theory also appear: Hall phase transitions (in a single layer) exhibit a selection rule $|pq'-p'q|=1$that also governs the Farey sequence. Indeed, plots of the semicircle law are also Farey diagrams.

In striped quantum Hall phases, the relationship is slightly more complex, because of the broken symmetry: $\begin{cases}\sigma \_{1}\sigma \_{2}+(\sigma \_{h}-\sigma \_{h}^{0})^{2}=(e^{2}/(2h))^{2}\\\sigma \_{h}^{0}=(N+1/2)e^{2}/h\end{cases}$Here *σ*1 and *σ*2 describe the macroscopic conductivity in directions aligned with and perpendicular to the stripes.